1. Introduction

   • Margaret R. Clarke

   Introduction Pages: 383 - 384

2. Population Characteristics of Howlers: Ecological Conditions or Group History

   • Colin A. Chapman
   • Sophia R. Balcomb

   OriginalPaper Pages: 385 - 403

3. Growth of Mantled Howler Groups in a Regenerating Costa Rican Dry Forest

   • Linda M. Fedigan
   • Lisa M. Rose
   • Rodrigo Morera Avila

   OriginalPaper Pages: 405 - 432

4. Agonistic and Affiliative Relationships of Adult Female Howlers (Alouatta palliata) in Costa Rica Over a 4-Year Period

   • Evan L. Zucker
   • Margaret R. Clarke

   OriginalPaper Pages: 433 - 449

5. Infant–Nonmother Interactions of Free-Ranging Mantled Howlers (Alouatta palliata) in Costa Rica

   • Margaret R. Clarke
   • Kenneth E. Glander
   • Evan L. Zucker

   OriginalPaper Pages: 451 - 472

6. Relation of Intergroup Variation in Allogrooming to Group Social Structure and Ectoparasite Loads in Red Howlers (Alouatta seniculus)

   • Marcelo R. Sánchez-Villagra
   • Theresa R. Pope
   • Viviana Salas

   OriginalPaper Pages: 473 - 491

7. Parasites of Wild Howlers (Alouatta spp.)

   • Michael Stuart
   • Vickie Pendergast
   • Margaret Clarke

   OriginalPaper Pages: 493 - 512

8. Physiological Ecology of Howlers (Alouatta): Energetic and Digestive Considerations and Comparison with the Colobinae

   • Katharine Milton

   OriginalPaper Pages: 513 - 548

9. Conservation Biology of the Genus Alouatta

   • Carolyn M. Crockett

   OriginalPaper Pages: 549 - 578

10. Effective Solutions for Howler Conservation

    • Robert H. Horwich

    OriginalPaper Pages: 579 - 598

11. Arboreal Quadrupedism and Forelimb Articular Anatomy of Red Howlers

    • Miguel A. Schön Ybarra

    OriginalPaper Pages: 599 - 613

12. Skeletal Pathologies in a Population of Alouatta palliata: Behavioral, Ecological, and Evolutionary Implications

    • David DeGusta
    • Katharine Milton

    OriginalPaper Pages: 615 - 650