1. Publisher's announcement

   Announcement Pages: i - i

2. Alternative formulations of multilevel selection

   • John Damuth
   • I. Lorraine Heisler

   OriginalPaper Pages: 407 - 430

3. The Why and How of Species

   • Ernst Mayr

   OriginalPaper Pages: 431 - 441

4. Haunt Me No Longer

   • Arthur L. Caplan
   • Walter J. Bock

   Continuing Commentary on Michael Ghiselin and Ernst Mayr (from volume 2 no. 2) Pages: 443 - 454

5. Species as individuals

   • Raphael Falk

   Continuing Commentary on Michael Ghiselin and Ernst Mayr (from volume 2 no. 2) Pages: 455 - 462

6. Discussion: Ghiselin and Mayr on species

   • Laurance Splitter

   Continuing Commentary on Michael Ghiselin and Ernst Mayr (from volume 2 no. 2) Pages: 462 - 467

7. The individuality thesis, essences, and laws of nature

   • Michael T. Ghiselin

   Continuing Commentary on Michael Ghiselin and Ernst Mayr (from volume 2 no. 2) Pages: 467 - 474

8. Can rationality and irrationality be reconciled?

   • Nils Chr Stenseth
   • Audfinn Tjønneland
   • Tore Lindholm

   Review and Responses Pages: 475 - 483

9. Reply from the author

   • Langdon Gilkey

   Review and Responses Pages: 485 - 495

10. A final note to Gilkey

    • Tore Lindholm
    • Nils Chr Stenseth
    • Audfinn Tjønneland

    Review and Responses Pages: 497 - 499

11. Letter to the Editor

    • John D. Collier

    Letter Pages: 501 - 503

12. Booknotes

    • M. R.

    BookReview Pages: 505 - 509

13. 1989 Summer Conference in history, philosophy, and social studies of biology

    Announcement Pages: 512 - 512