Abstract
In this chapter we redescribe Zangerlia dzamynchondi Sukhanov and Narmandakh 2006, a poorly described nanhsiungchelyid turtle from the Djadokhta Formation (Late Cretaceous, Campanian), Zamin Khond locality, southern Mongolia. We also present new observations on the holotype of Z. testudinimorpha, the type of the genus Zangerlia, and reassess some characters of this genus. Inclusion of these new data in a phylogenetic analyses of Adocusia (the clade uniting Adocidae and Nanhsiungchelyidae) allows us to reassess the phylogenetic position and relationships of Zangerlia. Our phylogenetic analysis does not support a monophyletic Zangerlia. All species of “Zangerlia” form a polytomy with Hanbogdemys orientalis, Anomalochelys angulata + Nanhsiungchelys wuchingensis, and Basilemys.
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Acknowledgments
We thank Magdalena Borsuk-Białynicka (ZPAL) for access to the holotype of Zangerlia testudinimorpha and her hospitality; Andrei Bochkov (Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia); and Walter Joyce, James Parham, and Ren Hirayama for their reviews of this paper. Jim Gardner and Walter Joyce deserve additional thanks for correcting the English in our manuscript. This study was undertaken with financial support by grants from the President of the Russian Federation to the Leading Scientific Schools (NSh-6560-2012.4) and from the Russian Foundation for Basic Research 08-05-00557-a and the Ministry of Education and Science of the Russian Federation.
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Appendices
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Appendix 1
Details about replacement character 27 used in phylogenetic analysis presented in this paper.
Description: Length of posterior plastral lobe: 0, more than distance between posterior ends of plastron and carapace; 1, less than distance between posterior ends of plastron and carapace.
Comments: This replaces character 27 in the previous analysis of Danilov and Syromyatnikova (2009). The primitive state of the new character 27 is found in the outgroups, adocids, and most nanhsiungchelyids. The derived condition is considered diagnostic for Zangerlia (Brinkman and Peng 1996; this chapter).
Codings: Xinjiangchelys levensis, 0; X. tianshanensis, 0; Carettochelys insculpta, 0; Apalone ferox, -; Yehguia tatsuensis, 0; Adocus aksary, ?; A. beatus/Adocus sp., 0; Ferganemys itemirensis, 0; F. verzilini, 0; Shachemys ancestralis, 0; S. baibolatica, 0; S. laosiana, 0; Kharakhutulia kalandadzei, 0; Zangerlia testudinimorpha, 1; Z. neimongolensis, 1; Z. ukhaachelys, ?; Z. dzamynchondi, 1; Hanbogdemys orientalis, 0; Anomalochelys angulata, ?; Nanhsiungchelys wuchingensis, ?; Basilemys variolosa, 0; B. nobilis, 0; B. sinuosa, 0; B. praeclara, 0.
Appendix 2
Characters coded for “Zangerlia” dzamynchondi and added to the matrix of Danilov and Syromyatnikova (2009).
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Danilov, I.G., Sukhanov, V.B., Syromyatnikova, E.V. (2013). Redescription of Zangerlia dzamynchondi (Testudines: Nanhsiungchelyidae) from the Late Cretaceous of Mongolia, with a Reassessment of the Phylogenetic Position and Relationships of Zangerlia . In: Brinkman, D., Holroyd, P., Gardner, J. (eds) Morphology and Evolution of Turtles. Vertebrate Paleobiology and Paleoanthropology. Springer, Dordrecht. https://doi.org/10.1007/978-94-007-4309-0_22
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