• K. Larsen
Part of the The Families and Genera of Vascular Plants book series (FAMILIES GENERA, volume 4)


Perennial, rhizomatous, non-aromatic herbs, terrestrial or more rarely epiphytic. Indumentum of uniseriate, multicellular or unicellular hairs. Stems terete, most often unbranched, leafy, sometimes spirally contorted; when branched, then secondary branches breaking through the leaf sheaths (Tapeinochilus). Leaves spirally arranged, sheaths closed, tubular; ligule present; petiole short; pulvinus absent; lamina narrowly to broadly elliptic, rolled up from one side to the other in bud. Inflorescence a strobilaceous spike, terminal on the leafy shoot or on a separate, short, leafless shoot, or flowers solitary, axillary (Monocostus). Bracts imbricate, with a linear, nectariferous callus below the tip, each subtending 1–2 flowers; bracteoles tubular or folded. Flowers epigynous, perfect, zygomorphic; calyx tubular, shortly 2–3-lobed; corolla 3-lobed, the lobes basally fused, imbricate in bud, unequal; stamen 1, often petaloid, bearing 2 2-sporangiate thecae, dehiscing introrsely by longitudinal slits, often appendaged at the apex. Labellum petaloid, staminodial, opposite to the stamen, as long as or much longer than the corolla, ± 3-lobed, often with crisped margin; basal part of stamen and labellum united into a papillate tube. Gynoecium 3(−2)-carpellate; ovary inferior, 3(−2)-Jocular; placentation axile; ovules numerous, anatropous, crassinucellar; septal nectaries 2, immersed in the apical part of the ovary; style filiform, commonly enclosed between the thecae; stigma Wet, with or without a dorsal appendage. Fruit a 3(−2)-locular capsule crowned by the persistent calyx, dehiscing loculicidally, rarely explosively, or indehiscent and irregularly breaking when old. Seeds numerous, aril white to yellow; operculum next to the prominent radicle; embryo straight; endosperm poorly developed; perisperm abundant, with copious starch in simple grains.


Leaf Axil Tall Herb Leafy Shoot Floral Tube Multicellular Hair 
These keywords were added by machine and not by the authors. This process is experimental and the keywords may be updated as the learning algorithm improves.


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Selected Bibliography

  1. Davis, G.L. 1966. See general references.Google Scholar
  2. Grootjen, C.J., Bouman, F. 1981. Development of the ovule and seed in Costus cuspidatus (N. E. Br.) Maas (Zingiberaceae), with special reference to the formation of the operculum. Bot. J. Linn. Soc. 83: 27-39.Google Scholar
  3. Hegnauer, R. 1986. See general references.Google Scholar
  4. Holttum, R.E. 1950. The Zingiberaceae of the Malay Peninsula. Gard. Bull. ( Singapore ) 13: 240-245.Google Scholar
  5. Johri, B.M. et al. 1992. See general references.Google Scholar
  6. Kato, M. 1996. Plant-pollinator interactions in the understory of a lowland mixed Dipterocarp forest in Sarawak. Am. J. Bot. 83: 732-743.Google Scholar
  7. Kirchoff, B.K. 1988. Inflorescence and flower development in Costus scaber. Can. J. Bot. 66: 339-345.Google Scholar
  8. Kress, W.J. 1990. The phylogeny and classification of the Zingiberales. Ann. Mo. Bot. Gard. 77: 698-721.Google Scholar
  9. Loesener, L.E.T. 1930. Zingiberaceae. Unterfam. II. Costoideae In: Engler, A., Prantl, K., Pflanzenfamilien, 2nd edn., 15a. Leipzig: W. Engelmann, pp. 628 - 640.Google Scholar
  10. Maas, P.J.M. 1972. Flora Neotropica, Monogr. 8. Costoideae (Zingiberaceae). New York: Hafner.Google Scholar
  11. Maas, P.J.M. 1977. Flora Neotropica, Mongr. 18. Renealimia (Zingiberaceae-Zingiberoideae), Costoideae (Additions). New York: The New York Botanical Garden.Google Scholar
  12. Pijl, L. van der 1954. Xylocopa and flowers in the tropics I-III. Proc. K. Ned. Akad. Wet. C 57: 413-423, 541 - 562.Google Scholar
  13. Punt, W. 1968. Pollen morphology of the American species of the subfamily Costoideae (Zingiberaceae). Rev. Palaeobot. Palynol. 7: 31-43.Google Scholar
  14. Schemske, D.W. 1980. The evolutionary significance of extrafloral nectar production by Costus woodsonii (Zingiberaceae): an experimental analysis of ant protection. J. Ecol. 68: 959 - 967.CrossRefGoogle Scholar
  15. Schemske, D.W. 1981. Floral convergence and pollinator sharing in two bee-pollinated, tropical herbs. Ecology 62: 946954.Google Scholar
  16. Schumann, K. 1904. Subfamilia II. Costoideae K. Schum. In: Engler, A. (ed.) Das Pflanzenreich, IV. 46. Leipzig, W. Engelmann, pp. 377 - 437.Google Scholar
  17. Stiles, F.G. 1978. Temporal organization of flowering among the hummingbird food plants of a tropical wet forest. Biotropica 10: 194 - 210.CrossRefGoogle Scholar
  18. Stone, D.E., Sellers, S.C., Kress, J.W. 1981. Ontogenetic and evolutionary implications of a neotenous exine in Tapeinochilos ( Zingiberales: Costaceae) pollen. Am. J. Bot. 68: 49-63.Google Scholar
  19. Takhtajan, A.L. 1982. See general references.Google Scholar
  20. Tillich, H.-J. 1995. Seedlings and systematics of the monocotyledons. In: Rudall, P.J., Cribb, P.J., Cutler, D.F., Humphries, C.J. (eds.) Monocotyledons: systematics and evolution, Vol. 1. London: Royal Botanic Gardens Kew, pp. 303 - 352.Google Scholar
  21. Tomlinson, P.B. 1969. See general references.Google Scholar
  22. Troll, W. 1928. Organisation und Gestalt im Bereich der Blüte. Berlin, Heidelberg, New York: Springer.Google Scholar
  23. Veh, R. von 1931. Untersuchungen und Betrachtungen zum Blattstelungsproblem. Flora 125: 83 - 154.Google Scholar

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© Springer-Verlag Berlin Heidelberg 1998

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  • K. Larsen

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