Abstract
Erect annual glabrous herbs, up to 1.5 m tall. Roots fibrous. Stems fistulose, fleshy or spongy, freely branching. Leaves alternate, simple, exstipulate, petiolate; lamina pinnately veined, margin entire. Inflorescence a dense terminal pedunculate acropetal conico-cylindric spike. Flowers small, tetracyclic, perfect, radially symmetrical, subtended by an inconspicuous spathulate bract and a pair of linear bracteoles. Calyx synsepalous, adnate to the ovary, forming a hypanthium; lobes 5, imbricate, persistent. Corolla sympetalous, radially symmetric, white or pale yellow, urceolate-campanulate, caducous; lobes 5, imbricate. Stamens 5, alternating with the corolla lobes; filaments distinct, adnate to the middle of the corolla tube, slightly dilated at base; anthers tetrasporangiate, dithecal, basifixed, distinct. Nectary 0. Gynoecium syncarpous, 2-locular; ovary half-inferior; ovules numerous; placentae axile, spongy; style simple, short or almost wanting; stigma capitate. Fruit a circumscissile capsule, enclosed by the accrescent calyx lobes. Seeds small, oblong, yellowish brown, shining, numerous, exarillate, testa striate; embryo straight; endosperm very scanty or lacking.
Access this chapter
Tax calculation will be finalised at checkout
Purchases are for personal use only
Selected Bibliography
Airy Shaw, H.K. 1948. Sphenocleaceae. In: van Steenis, C.G.G.J. (ed.) Flora Males. I, 4(1). Djakarta: Noordhoff-Kolff, pp. 27–28.
Airy Shaw, H.K. 1973. A dictionary of the flowering plants and ferns, ed. 8. Cambridge: Cambridge University Press.
d’Almeida, J.F.R. 1934. On the development of secondary aerating tissues in the stem of Sphenoclea zeylanica Gaertn. Proc. Indian Sci. Congr. Assoc. 1934: 314.
Bentham, G. 1876. Campanulaceae. In: Bentham, G., Hooker, J.D., Genera plantarum 2: 541–564. London: Reeve.
Cosner, M.E., Jansen, R.K., Lammers, T.G. 1994. Phylogenetic relationships in the Campanulales based on rbcL sequences. Plant Syst. Evol. 190: 79–95.
Cronquist, A. 1981. An integrated system of classification of flowering plants. New York: Columbia University Press.
Dahlgren, R.M.T. 1975. A system of classification of the angiosperms to be used to demonstrate the distribution of characters. Bot. Not. 128: 119–147.
Dahlgren, R.M.T. 1980. A revised system of classification of the angiosperms. Bot. J. Linn. Soc. 80: 91–124.
Dunbar, A. 1975. On pollen of Campanulaceae and related families with special reference to the surface ultrastructure II. Campanulaceae subfam. Cyphioideae and subfam. Lobelioideae; Goodeniaceae; Sphenocleaceae. Bot. Not. 128: 102–118.
Erbar, C. 1995. On the floral development of Sphenoclea zeylanica (Sphenocleaceae, Campanulales) – SEM investigations on herbarium material. Bot. Jahrb. Syst. 117: 469–493.
Gupta, D.P. 1959. Vascular anatomy of the flower of Sphenoclea zeylanica Gaertn. and some other related species. Proc. Natl. Inst. Sci. India 25B: 55–64.
Gustafsson, M.H.G., Bremer, K. 1995. Morphology and phylogenetic interrelationships of the Asteraceae, Calyceraceae, Campanulaceae, Goodeniaceae, and related families (Asterales). Amer. J. Bot. 82: 250–265.
Hiraia, N., Sakashita, S., Sano, T., Inoue, T., Ohigashi, H., Premasthira, C., Asakawa, Y., Harada, J., Fujii, Y. 2000. Allelochemicals of the tropical weed Sphenoclea zeylanica. Phytochemistry 55: 131–140.
Hutchinson, J. 1973. The families of flowering plants, ed. 3. Oxford: Clarendon Press.
Kausik, S.B., Subramanyam, K. 1946. A contribution to the life-history of Sphenoclea zeylanica Gaertn. Proc. Indian Acad. Sci. 23B: 274–280.
Lammers, T.G. 1992. Circumscription and phylogeny of the Campanulales. Ann. Missouri Bot. Gard. 79: 388–413.
Mohandas, C., Rao, Y.S., Sahu, S.C. 1981. Cultural control of rice root nematodes (Hirschmanniella spp.) with Sphenoclea zeylanica. Proc. Indian Acad. Sci., Animal Sci. 90: 373–376.
Monod, T. 1980. A propos du Sphenoclea zeylanica (Sphenocleaceae). Adansonia II, 20: 147–164.
Porembski, S., Koch, M. 1999. Inulin occurrence in Sphenoclea (Sphenocleaceae). Plant Biol. 1: 288–289.
Rosatti, T.J. 1986. The genera of Sphenocleaceae and Campanulaceae in the southeastern United States. J. Arnold Arb. 67: 1–64.
Schönland, S. 1889. Campanulaceae. In: Natürl. Pflanzenfamilien IV(5). Leipzig: W. Engelmann, pp. 40–70.
Soltis, D.E., Smith, S.A., Cellinese, N., Wurdack, K.J., Tank, D.C., Brockington, S.F., Refulio-Rodriguez, N.F., Walker, J.B., Moore, M.J., Carlsward, B.S., Bell, C.D., Latvis, M., Crawley, S., Black, C., Diouf, D., Xi, Z.X., Rushworth, C.A., Gitzendanner, M.A., Sytsma, K.J., Qiu, Y.L., Hilu, K.W., Davis, C.C., Sanderson, M.J., Beaman, R.S., Olmstead, R.G., Judd, W.S., Donoghue, M.J., Soltis, P.S. 2011. Angiosperm phylogeny: 17 genes, 640 taxa. Amer. J. Bot. 98: 704–730.
Subramanyam, K. 1950. A contribution to our knowledge of the systematic position of the Sphenocleaceae. Proc. Indian Acad. Sci. 31(B): 60–65.
Takhtajan, A. 1980. Outline of the classification of flowering plants (Magnoliophyta). Bot. Rev. 46: 225–359.
Takhtajan, A. 1981. Plant Life, vol. 5(2). Moskau: Proswjestschenije (in Russian).
Takhtajan, A. 1997. Diversity and classification of flowering plants. New York: Columbia University Press.
Thorne, R.F. 1976. A phylogenetic classification of the Angiospermae. In: Hecht, M.K., Steere, W.C., Wallace, B. (eds.) Evolutionary Biology 9. New York: Plenum Press, pp. 35–106.
Thorne, R.F. 1992. Classification and geography of the flowering plants. Bot. Rev. 58: 225–348.
Wagenitz, G. 1964. Reihe Campanulales. In: Melchior, H. (ed.) A. Engler’s Syllabus der Pflanzenfamilien, ed. 12, 2. Berlin: Bornträger, pp. 478–497.
Wilbur, R.L. 1976. Flora of Panama, Campanulaceae. Ann. Missouri Bot. Gard. 63: 593–655.
Author information
Authors and Affiliations
Corresponding author
Editor information
Editors and Affiliations
Rights and permissions
Copyright information
© 2016 Springer International Publishing Switzerland
About this chapter
Cite this chapter
Lammers, T.G. (2016). Sphenocleaceae. In: Kadereit, J., Bittrich, V. (eds) Flowering Plants. Eudicots. The Families and Genera of Vascular Plants, vol 14. Springer, Cham. https://doi.org/10.1007/978-3-319-28534-4_30
Download citation
DOI: https://doi.org/10.1007/978-3-319-28534-4_30
Published:
Publisher Name: Springer, Cham
Print ISBN: 978-3-319-28532-0
Online ISBN: 978-3-319-28534-4
eBook Packages: Biomedical and Life SciencesBiomedical and Life Sciences (R0)