Abstract
Annual or perennial herbs, subshrubs, shrubs, lianas or small trees; hermaphrodite or rarely unisexual and dioecious; indumentum of simple, unicellular, eglandular trichomes, sometimes with glandular cells. Leaves alternate, petiolate or sessile, exstipulate; lamina linear to suborbicular, margin usually entire, very rarely crenate or dentate, often revolute; apex mostly acute or acuminate; base cuneate or decurrent. Inflorescences thyrsoids, terminal or axillary; partial inflorescences scorpioid cymes. Flowers perfect, rarely unisexual, actinomorphic, chasmogamous, sessile or shortly pedicellate; calyx tube usually short, mostly campanulate, calyx lobes often unequal, linear to ovate, glabrous to densely hairy, mostly persistent after anthesis; corolla sympetalous, with subcircular to linear lobes; androecium haplostemonous, stamens antesepalous and borne on corolla tube, usually included, filaments short, linear; anthers dorsifixed, usually linear, tetrasporangiate; gynoecium superior, syncarpous, bicarpellate, usually 4-loculate with one ovule in each locule; ovule anatropous to hemitropous, unitegmic, tenuinucellate; style terminal with a conical stigmatic head with a basal ring-shaped stigma and a sterile, sometimes two-lobed apex; nectary disk at the base of the ovary. Fruit dry or fleshy, usually 4-seeded, rarely 1–2-seeded, falling apart into 1–4 mericarpids with 1–2 seeds each. Embryo minute, straight to curved, embedded in thin endosperm, cotyledons linear to ovoid.
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Diane, N., Hilger, H.H., Förther, H., Weigend, M., Luebert, F. (2016). Heliotropiaceae. In: Kadereit, J., Bittrich, V. (eds) Flowering Plants. Eudicots. The Families and Genera of Vascular Plants, vol 14. Springer, Cham. https://doi.org/10.1007/978-3-319-28534-4_17
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