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Heat, Color, and Flavor Compounds in Capsicum Fruit

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Book cover The Biological Activity of Phytochemicals

Part of the book series: Recent Advances in Phytochemistry ((RAPT,volume 41))

Abstract

Peppers are one of the oldest vegetables known and are popular in worldwide cuisine. Members of the Capsicum genus could be considered accomplished chemists, as these plants synthesize a variety of phytochemicals with valued roles in human health and nutrition as well as sensory perception. For thousands of years, humans have selected peppers for traits that affect heat, color, and flavor in the fruit. Heat in pepper fruit is attributed to production of capsaicinoids in the placenta; this alkaloid binds to nociceptive pain receptors in mammals. Color in pepper fruit is due to a combination of pigments: chlorophylls, carotenoids, and anthocyanins accumulating in the fruit wall or pericarp of the fruit resulting in green, yellow, or purple fruit at physiological immature stages and yellow, red, or orange fruit at mature stages. Flavor is the least characterized chemistry, but is in part the result of monoterpenoids and aliphatic aldehydes, which also accumulate in the fruit wall. The synthesis and accumulation of all these phytochemicals are dependent on the genetic background of the pepper, the developmental stage of the fruit, and environmental factors.

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References

  1. Perry L, Dickau R, Zarillo S, Holst I, Pearsall DM, Piperno DR, Berman MJ, Cooke RG, Rademaker K, Ranere AJ, Ramond JS, Sandweiss DJ, Scaramelli F, Tarble K, Zeidler JA (2007) Starch fossils and the domestication and dispersal of chili peppers (Capsicum spp. L.) in the Americas. Science 315:986–988

    Article  CAS  PubMed  Google Scholar 

  2. FAO (2006) Food and Agriculture Organization of the United States. http://www.fao.org

  3. USDA (2008) Vegetables and Melons Outlook. http://www.ers.usda.gov/publications/vgs/2008/08Aug/VGS328.pdf

  4. Pickersgill B (2007) Domestication of plants in the Americas: insights from Mendelian and molecular genetics. Ann Bot 100:925–940

    Article  PubMed  Google Scholar 

  5. Bosland PW (1996) Capsicums: Innovative uses of an ancient crop. In: Janick J (ed) Progress in new crops. ASHS Press, Arlington, VA, 479–487

    Google Scholar 

  6. Livingstone KD, Lackney VK, Blauth JR, Van Wijk R, Jahn MM (1999) Genome mapping in capsicum and the evolution of genome structure in the Solanaceae. Genetics 152:1183–1202

    CAS  PubMed  Google Scholar 

  7. Wu F, Eannetta NT, Xu Y, Durrett R, Mazourek M, Jahn MM, Tanksley SD (2009) A COSII genetic map of the pepper genome provides a detailed picture of synteny with tomato and new insights into recent chromosome evolution in the genus Capsicum. Theor Appl Genet 118:1279–1293

    Article  CAS  PubMed  Google Scholar 

  8. Paran I, Van Der Knaap E (2007) Genetic and molecular regulation of fruit and plant domestication traits in tomato and pepper. J Exp Bot 58:3841–3852

    Article  CAS  PubMed  Google Scholar 

  9. Wang D, Bosland P (2006) The genes of Capsicum. HortScience 41:1169–1187

    CAS  Google Scholar 

  10. DellaPenna D, Pogson BJ (2006) Vitamin synthesis in plants: tocopherols and carotenoids. Annu Rev Plant Biol 57:711–738

    Article  CAS  PubMed  Google Scholar 

  11. Deepa N, Kaur C, George B, Singh B, Kapoor HC (2005) Antioxidant constituents in some sweet pepper (Capsicum annuum L.) genotypes during maturity. Food Sci Technol 40:121–129

    Google Scholar 

  12. Gross J (1991) Pigments in vegetables: chlorophylls and carotenoids. Van Nostand Reinhold, New York, NY

    Google Scholar 

  13. Sergio AR, Paiva MD, Rusell RM (1999) β-carotene and other carotenoids as antioxidants. J Amer Coll Nutr 18:426–433

    Google Scholar 

  14. Schemske DW, Bradshaw HD (1999) Pollinator preference and the evolution of floral traits in monkeyflowers (Mimulus). Proc Natl Acad Sci USA 96:11910–11915

    Article  CAS  PubMed  Google Scholar 

  15. Wang HC, Huang XM, Hu GB, Yang Z, Huang HB (2005) A comparative study of chlorophyll loss and its related mechanism during fruit maturation in the pericarp of fast- and slow-degreening litchi pericarp. Sci Hortic 106:247–257

    Article  CAS  Google Scholar 

  16. Smith PG (1950) Inheritance of brown and green mature color in peppers. J Hered 41:138–140

    CAS  PubMed  Google Scholar 

  17. Lightbourn GJ, Griesbach RJ, Novotny JA, Clevidence BA, Rao DD, Stommel JR (2008) Effects of anthocyanin and carotenoid combinations on foliage and immature fruit color of Capsicum annuum L. J Hered 99:105–111

    Article  CAS  PubMed  Google Scholar 

  18. Deli J, Molnar P, Toth G (2001) Carotenoid composition in the fruits of red paprika (Capsicum annuum var. lycopersiciforme rubrum) during ripening; biosynthesis of carotenoids in red paprika. J Agric Food Chem 49:1517–1523

    Article  CAS  PubMed  Google Scholar 

  19. Matsufuji H, Nakamura H, Chino M, Takeda M (1998) Antioxidant activity of capsanthin and the fatty acid esters in paprika (Capsicum annuum). J Agric Food Chem 46:3468–3472

    Article  CAS  Google Scholar 

  20. Wall MM, Waddell CA, Bosland PW (2001) Variation in β-carotene and total carotenoid content in fruits of Capsicum. HortScience 36:746–749

    CAS  Google Scholar 

  21. Auldrige ME, McCarty DR, Klee HJ (2006) Plant carotenoid cleavage oxygenases and their apocarotenoid products. Curr Opin Plant Biol 9:315–321

    Article  Google Scholar 

  22. von Lintig J, Vogt K (2004) Vitamin A formation in animals: molecular identification and functional characterization of carotenoid cleaving enzymes. J Nutr 134:251S-256S

    Google Scholar 

  23. USDA (2000) Nutrition and Your Health: Dietary Guidelines for Americans. http://www.cnpp.usda.gov/Publications/DietaryGuidelines/2000/2000DGProfessionalBooklet.pdf.

  24. Purseglove J, Brown E, Green C, Robbins S (1981) Spices. Longman, London

    Google Scholar 

  25. Hugueney P, Badillo A, Chen H-C, Klein A, Hirschberg J, Camara B, Kuntz M (1995) Metabolism of cyclic carotenoids: a model for the alteration of this biosynthetic pathway in Capsicum annuum chromoplasts. Plant J 8:417–424

    Article  CAS  PubMed  Google Scholar 

  26. Fraser P, Bramley P (2004) The biosynthesis and nutritional uses of carotenoids. Prog Lipid Res 43:228–265

    Article  CAS  PubMed  Google Scholar 

  27. Hugueney P, Bouvier F, Badillo A, Quennemet J, d'Harlingue A, Camara B (1996) Developmental and stress regulation of gene expression for plastid and cytosolic isoprenoid pathways in pepper fruits. Plant Physiol 111:619–626

    Article  CAS  PubMed  Google Scholar 

  28. Cunningham FX, Grantt E (1998) Genes and enzymes of carotenoid biosynthesis in plants. Annu Rev Plant Physiol Plant Mol Biol 49:557–583

    Article  CAS  PubMed  Google Scholar 

  29. Romer S, Hugueney P, Bouvier F, Camara B, Kuntz M (1993) Expression of the genes encoding the early carotenoid biosynthetic enzymes in Capsicum annuum. Biochem Biophys Res Commun 196:1414–1421

    Article  CAS  PubMed  Google Scholar 

  30. Bouvier F, Keller Y, d'Harlingue A, Camara B (1998) Xanthophyll biosynthesis: molecular and functional characterization of carotenoid hydroxlases from pepper fruits (Capscium annuum L.). Biochim Biophys Acta 1391:320–328

    CAS  PubMed  Google Scholar 

  31. Bouvier F, d'Harlingue A, Hugueney P, Marin E, Marion-Poll A, Camara B (1996) Xanthophyll biosynthesis: cloning, expression, functional reconstitution, and regulation of β-cyclohexenyl carotenoid epoxidase from pepper (Capsicum annuum). J Biol Chem 271:28861–22867

    Google Scholar 

  32. Bouvier F, Hugueney P, d'Harlingue A, Kuntz M, Camara B (1994) Xanthophyll biosynthesis in chromoplasts: isolation and molecular cloning of an enzyme catalyzing the conversion of 5,6-epoxycarotenoid into ketocarotenoid. Plant J 6:45–54

    Article  CAS  PubMed  Google Scholar 

  33. Hurtado-Hernandez H, Smith PG (1985) Inheritance of mature fruit color in Capsicum annuum L. J Hered 76:211–213

    Google Scholar 

  34. Lefebvre V, Kuntz M, Camara B, Palloix A (1998) The capsanthin-capsorubin synthase gene: a candidate gene for the y locus controlling the red fruit colour in pepper. Plant Mol Biol 36:785–789

    Article  CAS  PubMed  Google Scholar 

  35. Huh JH, Kan BC, Nahm SH, Kim S, Ha KS, Lee MH, Kim BD (2001) A candidate gene approach identified phytoene synthase as the locus for mature fruit color in red pepper (Capsicum spp.). Theor Appl Genet 102:524–530

    Article  CAS  Google Scholar 

  36. Guzman I, Hamby S, Romero J, Bosland PW, O’Connell MA (2010) Variability of Carotenoid Biosynthesis in Orange Colored Capsicum spp. Plant Science, 179:49–59

    Article  CAS  PubMed  Google Scholar 

  37. Lang YQ, Yanagawa S, Sasanuma T, Sasakuma T (2004) Orange fruit color in Capsicum due to deletion of capsanthin-capsorubin synthesis gene. Breed Sci 54:33–39

    Article  CAS  Google Scholar 

  38. Popovsky S, Paran I (2000) Molecular genetics of the y locus in pepper: its relation to capsanthin-capsorubin synthase and to fruit color. Theor Appl Genet 101:86–89

    Article  CAS  Google Scholar 

  39. Thorup TA, Tanyolac B, Livingstone KD, Popovsky S, Paran I, Jahn M (2000) Candidate gene analysis of organ pigmentation loci in the Solanaceae. Proc Natl Acad Sci USA 97:11192–11197

    Article  CAS  PubMed  Google Scholar 

  40. Hornero-Mendez D, Minguez-Mosquera MI (2000) Xanthophyll esterification accompanying carotenoid overaccumulation in chromoplasts of Capsicum annuum ripening fruits is a constitutive process and useful for ripeness index. J Agric Food Chem 48:1617–1622

    Article  CAS  PubMed  Google Scholar 

  41. Ye X, Al-Babili S, Kloti A, Zhang J, Lucca P, Beyer P, Potrykus I (2000) Engineering provitamin A (β-carotene) biosynthesis pathway into (carotenoid-free) rice endosperm. Science 287:303–305

    Article  CAS  PubMed  Google Scholar 

  42. Paran I, Van Der Voort JR, Lefebvre V, Jahn MM, Landry L, Van Schriek M, Tanyolac B, Caranta C, Chaim AB, Livingstone K, Palloix A, Peleman J (2004) An integrated genetic linkage map of pepper (Capsicum spp.). Mol Breed 13:251–261

    Article  CAS  Google Scholar 

  43. Daskalov S, Poulos JM (1994) Updated Capsicum gene list. Capsicum Eggplant Newsl 13:15–26

    Google Scholar 

  44. Ben Chaim A, Paran I, Grube R, Jahn MM, Van Wijk R, Peleman J (2001) QTL mapping of fruit related traits in pepper (Capsicum annuum). Theor Appl Genet 102:1016–1028

    Article  Google Scholar 

  45. Borovsky Y, Oren-Shamir M, Ovadia R, De Jong W, Paran I (2004) The A locus that controls anthocyanin accumulation in pepper encodes a MYB transcription factor homologous to Anthocyanin2 of Petunia. Theor Appl Genet 109:23–29

    Article  CAS  PubMed  Google Scholar 

  46. Billing J, Sherman PW (1998) Antimicrobial functions of spices: why some like it hot. Quart Rev Biol 73:3–49

    Article  CAS  PubMed  Google Scholar 

  47. Govindarajan V, Sathyanarayana M (1991) Capsicum - production, technology, chemistry, and quality. Part V - Impact on physiology, pharmacology, nutrition and metabolism; structure, pungency, pain and desensitization sequences. Crit Rev Food Sci Nutr 29:435–474

    Article  CAS  PubMed  Google Scholar 

  48. Winter J, Bevan S, Campbell EA (1995) Capsaicin and pain mechanisms. Brit J Anaesth 75:157–168

    CAS  PubMed  Google Scholar 

  49. Caterina MJ, Schumacher MA, Tominaga M, Rosen TA, Levine JD, Julius D (1997) The capsaicin receptor: a heat-activated ion channel in pain pathway. Nature 389:816–824

    Article  CAS  PubMed  Google Scholar 

  50. Tominaga M, Caterina MJ, Malmberg AB, Rosen TA, Gilbert H, Skinner K, Raumann BE, Basbaum AI, Julius D (1998) The cloned capsaicin receptor integrates multiple pain-producing stimuli. Neuron 21:531–543

    Article  CAS  PubMed  Google Scholar 

  51. Bennett DJ, Kirby GW (1968) Constitution and biosynthesis of capsaicin. J Chem Soc C:442–446

    Google Scholar 

  52. Leete E, Louden M (1968) Biosynthesis of capsaicin and dihydrocapsaicin in Capsicum frutescens. J Am Chem Soc 90:6837–6841

    Article  CAS  PubMed  Google Scholar 

  53. Krajewska A, Powers J (1988) Sensory properties of naturally occurring capsaicinoids. J Food Sci 53:902–905

    Article  Google Scholar 

  54. Collins MD, Mayer-Wasmund L, Bosland PW (1995) Improved method for quantifying capsaicinoids in Capsicum using high performance liquid chromatography. HortScience 30:137–139

    CAS  Google Scholar 

  55. Tewksbury JJ, Nabhan GP (2001) Directed deterrence by capsaicin in chillies. Nature 412:403–404

    Article  CAS  PubMed  Google Scholar 

  56. Tewksbury J, Reagan K, Machnicki N, Carlo T, Haak D, Calderon-Penaloza A, Levey D (2008) Evolutionary ecology of pungency in wild chilies. Proc Natl Acad Sci USA 105:11808–11811

    Article  CAS  PubMed  Google Scholar 

  57. Fujiwake H, Suzuki T, Iwai K (1982) Capsaicinoid formation in the protoplast from the placenta of Capsicum fruits. Agric Biol Chem 46:2591–2592

    CAS  Google Scholar 

  58. Fujiwake H, Suzuki T, Iwai K (1982) Intracellular distributions of enzymes and intermediates involved in biosynthesis of capsaicin and its analogues in Capsicum fruits. Agric Biol Chem 46:2685–2689

    CAS  Google Scholar 

  59. Hall R, Holden M, Yeoman M (1987) The accumulation of phenylpropanoid and capsaicinoid compounds in cell cultures and whole fruit of the chili pepper, Capsicum frutescens Mill. Plant Cell Tissue Organ Cult 8:163–176

    Article  CAS  Google Scholar 

  60. Iwai K, Suzuki T, Fujiwake H (1979) Formation and accumulation of pungent principle of hot pepper fruits, capsaicin and its analogues in Capsicum annuum var. annuum cv. Karayatsubusa at different growth stages after flowering. Agric Biol Chem 43:2493–2498

    CAS  Google Scholar 

  61. Ochoa-Alejo N, Gomez-Peralta JE (1993) Activity of enzymes involved in capsaicin biosynthesis in callus tissue and fruits of chili pepper (Capsicum annuum L.). J Plant Physiol 141:147–152

    CAS  Google Scholar 

  62. Sukrasno N, Yeoman MM (1993) Phenylpropanoid metabolism during growth and development of Capsicum frutescens fruits. Phytochemistry 32:839–844

    Article  CAS  Google Scholar 

  63. Suzuki T, Kawada T, Iwai K (1981) The precursors affecting the composition of capsaicin and its analogues in the fruits of Capsicum annuum var. annuum cv. Karayatsubusa. Agric Biol Chem 45:535–537

    CAS  Google Scholar 

  64. Aluru MR, Mazourek M, Landry LG, Curry J, Jahn MM, O'Connell MA (2003) Differential expression of fatty acid synthase genes, Acl, Fat and Kas in Capsicum fruit. J Exp Bot 54:1655–1664

    Article  CAS  PubMed  Google Scholar 

  65. Curry J, Aluru M, Mendoza M, Nevarez J, Melendrez M, O'Connell MA (1999) Transcripts for capsaicinoid biosynthetic genes are differentially accumulated in pungent and non-pungent Capsicum spp. Plant Sci 148:47–57

    Article  CAS  Google Scholar 

  66. Kim M, Kim S, Kim S, Kim B-D (2001) Isolation of cDNA clones differentially accumulated in the placenta of pungent pepper by suppression subtractive hybridization. Mol Cells 11:213–219

    CAS  PubMed  Google Scholar 

  67. Mazourek M, Pujar A, Borovsky Y, Paran I, Mueller L, Jahn MM (2009) A dynamic interface for capsaicinoid systems biology. Plant Physiol 150:1806–1821

    Article  CAS  PubMed  Google Scholar 

  68. Orlova I, Marshall-Colon A, Schnepp J, Wood B, Varbanova M, Fridman E, Blakeslee JJ, Peer WA, Murphy AS, Rhodes D, Pichersky E, Dudareva N (2006) Reduction of benzoid synthesis in petunia flowers reveals multiple pathways to benzoic acid and enhancement in auxin transport. Plant Cell 18:3458–3475

    Article  CAS  PubMed  Google Scholar 

  69. Thiele R, Mueller-Seitz E, Petz M (2008) Chili pepper fruits: presumed precursors of fatty acids characteristic for capsaicinoids. J Agric Food Chem 56:4219–4224

    Article  CAS  PubMed  Google Scholar 

  70. Fujiwake H, Suzuki T, Oka S, Iwai K (1980) Enzymatic formation of capsaicinoid from vanillylamine and iso-type fatty acids by cell-free extracts of Capsicum annuum var. annuum cv. Karayatsubusa. Agric Biol Chem 44:2907–2912

    CAS  Google Scholar 

  71. Stewart C, Kang B-C, Liu K, Mazourek M, Moore SL, Yoo EY, Kim B-D, Paran I, Jahn MM (2005) The Pun1 gene for pungency in pepper encodes a putative acyltransferase. Plant J 42:675–688

    Article  CAS  PubMed  Google Scholar 

  72. Stewart C, Mazourek M, Stellari GM, O’Connell MA, Jahn M (2007) Genetic control of pungency in C. chinense via the Pun 1 locus. J Exp Bot 58:979–991

    Article  CAS  PubMed  Google Scholar 

  73. Suzuki T, Fujiwake H, Iwai K (1980) Intracellular localization of capsaicin and its analogues, capsaicinoid, in Capsicum fruit. 1. Microscopic investigation of the structure of the placenta of Capsicum annuum var. annuum cv. Karayatsubusa. Plant Cell Physiol 21:839–853

    CAS  Google Scholar 

  74. Ohta Y (1963) Physiological and genetical studies on the pungency of Capsicum IV. Secretory organ, receptacles and distribution of capsaicin in the Capsicum fruits. Jap J Breed 12:179–183

    Google Scholar 

  75. Heiser CB, Smith PG (1953) The cultivated Capsicum peppers. Econ Bot 7:214–227

    Article  Google Scholar 

  76. Blum E, Liu K, Mazourek M, Yoo EY, Jahn MM, Paran I (2002) Molecular mapping of the C locus for presence of pungency in Capsicum. Genome 45:702–705

    Article  CAS  PubMed  Google Scholar 

  77. Blum E, Mazourek M, O’Connell MA, Curry J, Thorup T, Liu K, Jahn MM, Paran I (2003) Molecular mapping of capsaiciniod biosynthesis genes and QTL analysis for capsaicinoid content in Capsicum. Theor Appl Genet 108:79–86

    Article  CAS  PubMed  Google Scholar 

  78. Eshbaugh WH (1980) The taxonomy of the genus Capsicum (Solanaceae). Phytologia 47:153–166

    Google Scholar 

  79. Tewksbury J, Manchego C, Haak D, Levey D (2006) Where did the chili get its spice? Biogeography of capsiaicinoid production in ancestral wild chili species. J Chem Ecol 32:547–564

    Article  CAS  PubMed  Google Scholar 

  80. Iwai K, Lee K-R, Kobashi M, Suzuki T (1977) Formation of pungent principles in fruits of sweet pepper, Capsicum annuum L. var. grossum during post-harvest ripening under continuous light. Agric Biol Chem 41:1873–1876

    CAS  Google Scholar 

  81. Iwai K, Suzuki T, Lee K-R, Kobashi M, Oka S (1977) In vivo and in vitro formation of dihydrocapsaicin in sweet pepper fruits, Capsicum annuum L. var. grossum. Agric Biol Chem 41:1877–1882

    CAS  Google Scholar 

  82. Votava E, Bosland P (2002) Novel sources of non-pungency in Capsicum species. Capsicum Eggplant Newslett 21:66–68

    Google Scholar 

  83. Garces-Claver A, Gil-Ortega R, Alvarez-Fernandez A, Arnedo-Andres M (2007) Inheritance of capsaicin and dihydrocapsaicin, determined by HPLC-ESI/MS, in an intraspecific cross of Capsicum annuum L. J Agric Food Chem 55:6951–6957

    Article  CAS  PubMed  Google Scholar 

  84. Zewdie Y, Bosland PW (2000) Capsaicinoid inheritance in an interspecific hybridization of Capsicum annuum x C. chinense. HortScience 125:448–453

    CAS  Google Scholar 

  85. Ben Chaim A, Borovsky Y, Falise M, Mazourek M, Kung BC, Paran I, Jahn MM (2006) QTL analysis for capsaicinoid content in Capsicum. Theor Appl Genet 113:1481–1490

    Article  CAS  PubMed  Google Scholar 

  86. Harvell K, Bosland PW (1997) The environment produces a significant effect on pungency of chiles. HortScience 32:1292

    Google Scholar 

  87. Zewdie Y, Bosland PW (2000) Evaluation of genotype, environment, and genotype-by-environment interaction for capsaicinoids in Capsicum annuum L. Euphytica 111:185–190

    Article  CAS  Google Scholar 

  88. Mueller-Seitz E, Hiepler C, Petz M (2008) Chili pepper fruits: content and pattern of capsaicinoids in single fruits of different ages. J Agric Food Chem 56:12114–12121

    Article  CAS  PubMed  Google Scholar 

  89. Zewdie Y, Bosland PW (2000) Pungency of chile (Capsicum annuum L.) fruit is affected by node position. HortScience 35:1174

    Google Scholar 

  90. Buttery RG, Seifert RM, Guadagni DG, Ling LC (1969) Characterization of some volatile constituents of bell peppers. J Agric Food Chem 17:1322–1327

    Article  Google Scholar 

  91. Zimmermann M, Schieberle P (2000) Important odorants of sweet bell pepper powder (Capsicum annuum cv. annuum): differences between samples of Hungarian and Moroccan origin. Eur Food Res Technol 211:175–180

    Article  CAS  Google Scholar 

  92. Mazidaa MM, Sallehb MM, Osman H (2005) Analysis of volatile aroma compounds of fresh chilli (Capsicum annuum) during stages of maturity using solid phase microextraction (SPME). J Food Comp Anal 18:427–437

    Google Scholar 

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Acknowledgment

This work was supported in part by NM Agricultural Experiment Station, NIH grants GM08136 and GM61222, and USDA grant CSREES 2009-34604-19939.

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Correspondence to Mary A. O’Connell .

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Guzman, I., Bosland, P.W., O’Connell, M.A. (2011). Heat, Color, and Flavor Compounds in Capsicum Fruit. In: Gang, D. (eds) The Biological Activity of Phytochemicals. Recent Advances in Phytochemistry, vol 41. Springer, New York, NY. https://doi.org/10.1007/978-1-4419-7299-6_8

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