Complications of Inheritance
Three papers were published within a few weeks of each other in 1900 in which the authors—de Vries, Correns, and Tschermak— brought Mendel’s remarkable paper back to the attention of the scientific world and confirmed and extended his results. Segregation and random recombination were unquestionably facts. In 1906, however, Bateson and Punnett discovered evidence indicating that recombination was not universal. In our discussion of complementary gene action in Chapter 2, we saw that in their cross between white-flowered sweet peas the F1 was purple-flowered because complementary factors C and R were brought together in it, and a factor B modified the color from red to purple by the addition of blue. The original reason for this particular cross was to study inheritance of pollen shape; the color problem just presented itself by chance, as problems seem to do in scientific research. But what did they learn about inheritance of pollen-grain shape?
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