© 2018

Water Bears: The Biology of Tardigrades

  • Ralph O. Schill

Part of the Zoological Monographs book series (ZM, volume 2)

Table of contents

  1. Front Matter
    Pages i-viii
  2. Nadja Møbjerg, Aslak Jørgensen, Reinhardt Møbjerg Kristensen, Ricardo C. Neves
    Pages 57-94
  3. Aslak Jørgensen, Reinhardt M. Kristensen, Nadja Møbjerg
    Pages 95-114
  4. S. J. McInnes, P. J. A. Pugh
    Pages 115-129
  5. Roberto Guidetti, Roberto Bertolani
    Pages 131-143
  6. Roberto Bertolani, Lorena Rebecchi
    Pages 145-161
  7. Diane R. Nelson, Paul J. Bartels, Noemi Guil
    Pages 163-210
  8. Tiziana Altiero, Atsushi C. Suzuki, Lorena Rebecchi
    Pages 211-247
  9. Roberto Guidetti, Nadja Møbjerg
    Pages 249-271
  10. Ralph O. Schill, Steffen Hengherr
    Pages 273-293
  11. Steffen Hengherr, Ralph O. Schill
    Pages 295-310
  12. K. Ingemar Jönsson, Eliana B. Levine, Andrzej Wojcik, Siamak Haghdoost, Mats Harms-Ringdahl
    Pages 311-330
  13. Thomas C. Boothby
    Pages 331-347
  14. Peter Degma
    Pages 349-369
  15. Peter Degma, Roberto Guidetti
    Pages 371-409
  16. Back Matter
    Pages 411-419

About this book


Offering extensive information on tardigrades, this volume begins with a chapter on the history of tardigrades, from the first description by Goeze in 1773, until 1929, when the most comprehensive monographic approach by E. Marcus was published. Tardigrades’ organ systems, including their integument, body cavity, digestive, muscular, nervous and reproductive systems, as well as their overall external morphology, are summarized in the second chapter. Subsequent chapters present the current state of knowledge on tardigrade phylogeny, biogeography, paleontology, cytology and cytogenetics. In addition, the book provides insights into the ecology of tardigrades in marine, freshwater and terrestrial habitats. The reproduction, development and life cycles are summarized and the extraordinary environmental adaptations of encystment and cyclomorphosis, desiccation tolerance, freezing tolerance and radiation tolerance are discussed in detail. Further chapters provide an overview of key approaches in molecular tardigrade studies and describe techniques for sampling and sample processing. The book closes with a list of tardigrade taxa up to a sub-generic level, including the type species of each genus, the numbers of lower taxa in each taxon, and the main environments in which the taxa were found.
Given its depth of coverage, the volume offers an invaluable resource for scientists from various disciplines who plan to research tardigrades, and for all others who are interested in these fascinating animals.


Tardigrada biodiversity anhydrobiosis cryobiosis cryptobiosis diapause dormancy dormant state eutardigrades heat tolerance heterotardigrades extreme environments parthenogentic

Editors and affiliations

  • Ralph O. Schill
    • 1
  1. 1.Institute of Biological Materials and Biomolecular SystemsUniversity of StuttgartStuttgartGermany

About the editors

​Ralph O. Schill
University of Stuttgart
Institute of Biological Materials and Biomolecular Systems
Stuttgart, Germany

Bibliographic information

  • Book Title Water Bears: The Biology of Tardigrades
  • Editors Ralph O. Schill
  • Series Title Zoological Monographs
  • Series Abbreviated Title Zoological Monographs
  • DOI
  • Copyright Information Springer Nature Switzerland AG 2018
  • Publisher Name Springer, Cham
  • eBook Packages Biomedical and Life Sciences Biomedical and Life Sciences (R0)
  • Hardcover ISBN 978-3-319-95701-2
  • eBook ISBN 978-3-319-95702-9
  • Series ISSN 2523-3904
  • Series E-ISSN 2523-3912
  • Edition Number 1
  • Number of Pages VIII, 419
  • Number of Illustrations 61 b/w illustrations, 57 illustrations in colour
  • Topics Zoology
    Evolutionary Biology
    Animal Ecology
  • Buy this book on publisher's site