Table 1 Number of ground beetles (Carabidae) in field margin habitats and fields over 2006–2007

From: Ground beetles (Carabidae) of field margin habitats

Species K PI R1 L PII R2 Total Mean no. in the fields Mean no. in the field margins
Acupalpus meridianus (Linnaeus, 1761) 1 6 4 1 0 2 14 3.0 2.0
Acupalpus mixtus (Herbst, 1784) 13 0 1 0 0 1 15 0.0 3.8
Agonum gracile* (Sturm, 1824) 0 0 0 1 0 0 1 0.0 0.3
Agonum sexpunctatum* L* (Linnaeus, 1758) 0 0 0 1 0 0 1 0.0 0.3
Agonum viduum* (Panzer, 1796) 19 0 4 1 0 7 31 0.0 7.8
Amara aenea (De Geer, 1774) 7 8 11 1 0 2 29 4.0 5.3
Amara apricaria* (Paykull, 1790) 9 0 6 2 2 0 19 1.0 4.3
Amara aulica* (Panzer, 1796) 8 1 7 35 2 0 53 1.5 12.5
Amara bifrons (Gyllenhal, 1810) 15 0 13 2 7 10 47 3.5 10.0
Amara communis (Panzer, 1797) 1 0 0 0 0 0 1 0.0 0.3
Amara convexiuscula (Marsham, 1802) 0 0 0 1 0 0 1 0.0 0.3
Amara curta Dejean, 1828 0 0 0 1 0 0 1 0.0 0.3
Amara familiaris (Duftschmid, 1812) 40 2 12 22 1 5 82 1.5 19.8
Amara fulva (O.F. Müller, 1776) 0 0 1 0 0 0 1 0.0 0.3
Amara plebeja* (Gyllenhal, 1810) 1 7 1 2 1 8 20 4.0 3.0
Amara ovata (Fabricius, 1792) 8 3 16 10 1 6 44 2.0 10.0
Amara similata (Gyllenhal, 1810) 3 1 1 0 1 1 7 1.0 1.3
Amara spreta Dejean, 1831 0 1 0 0 0 0 1 0.5 0.0
Amara sp. Bonelli, 1810 3 7 4 0 0 3 17 3.5 2.5
Anchomenus dorsalis* (Pontoppidan, 1763) 521 195 343 172 111 129 1471 153.0 291.3
Anisodactylus binotatus (Fabricius, 1787) 0 0 0 0 0 2 2 0.0 0.5
Asaphidion flavipes (Linnaeus, 1761) 4 8 2 16 0 4 34 4.0 6.5
Badister bipustulatus (Fabricius, 1792) 5 0 9 21 0 10 45 0.0 11.3
Bembidion femoratum* Sturm, 1825 3 1 4 5 8 3 24 4.5 3.8
Bembidion lampros* (Herbst, 1784) 23 28 63 22 1 12 149 14.5 30.0
Bembidion properans* (Stephens, 1828) 63 233 438 32 42 68 876 137.5 150.3
Bembidion quadrimaculatum* (Linnaeus, 1761) 2 77 26 3 52 3 163 64.5 8.5
Bembidion ustulatum* (Linnaeus, 1758) 14 99 12 76 72 32 305 85.5 33.5
Bembidion sp.* Latreille, 1802 0 3 0 11 3 0 17 3.0 2.8
Broscus cephalotes (Linnaeus, 1758) 0 4 3 0 1 0 8 2.5 0.8
Calathus ambiguus* (Paykull, 1790) 256 80 129 33 61 20 579 70.5 109.5
Calathus erratus* (C.R. Sahlberg, 1827) 15 10 9 1 10 2 47 10.0 6.8
Calathus fuscipes* Goeze, 1777 88 20 41 3 3 15 170 11.5 36.8
Calathus melanocephalus* (Linnaeus, 1758) 10 5 2 26 46 10 99 25.5 12.0
Calathus micropterus* (Duftschmid, 1812) 2 0 2 3 0 0 7 0.0 1.8
Calosoma auropunctatum (Herbst, 1784) 0 0 1 0 0 0 1 0.0 0.3
Carabus auratus Linnaeus, 1761 224 147 333 85 7 158 954 77.0 200.0
Carabus cancellatus Illiger, 1798 89 68 243 77 15 39 531 41.5 112.0
Carabus convexus Fabricius, 1775 23 0 1 11 0 2 37 0.0 9.3
Carabus hortensis Linnaeus, 1758 1 0 0 11 0 1 13 0.0 3.3
Carabus granulatus Linnaeus, 1758 2 0 0 0 0 0 2 0.0 0.5
Carabus nemoralis O.F. Müller, 1764 5 0 3 23 0 0 31 0.0 7.8
Carabus violaceus Linnaeus, 1758 0 0 0 1 0 0 1 0.0 0.3
Chlaenius nigricornis (Fabricius, 1787) 1 1 1 0 0 6 9 0.5 2.0
Clivina collaris (Herbst, 1784) 0 21 1 13 49 6 90 35.0 5.0
Clivina fossor Linnaeus, 1758 0 1 0 0 4 1 6 2.5 0.3
Dyschirus globosus (Herbst, 1784) 0 1 0 1 0 0 2 0.5 0.3
Elaphrus riparius Linnaeus, 1758 0 0 1 1 0 0 2 0.0 0.5
Harpalus affinis* (Schrank, 1781) 149 131 283 78 166 217 1024 148.5 181.8
Harpalus anxius (Duftschmid, 1812) 0 0 0 0 1 0 1 0.5 0.0
Harpalus brevicollis Audinet-Serville, 1821 3 3 5 4 3 3 21 3.0 3.8
Harpalus latus (Linnaeus, 1758) 0 0 0 4 0 1 5 0.0 1.3
Harpalus rubripes (Duftschmid, 1812) 0 0 9 2 0 16 27 0.0 6.8
Harpalus signaticornis (Duftschmid, 1812) 1 0 1 0 2 0 4 1.0 0.5
Harpalus smaragdinus (Duftschmid, 1812) 0 0 1 1 0 0 2 0.0 0.5
Harpalus tardus (Panzer, 1796) 67 3 25 8 2 12 117 2.5 28.0
Harpalus vernalis (Fabricius, 1801) 2 0 1 1 0 3 7 0.0 1.8
Harpalus sp. Latreille, 1802 4 0 1 4 0 9 18 0.0 4.5
Leistus ferrugineus (Linnaeus, 1758) 29 8 15 19 0 3 74 4.0 16.5
Limodromus assimilis* (Pontoppidan, 1763) 1190 6 3 1028 15 6 2248 10.5 556.8
Loricera pilicornis* (Fabricius, 1775) 31 63 18 22 25 21 180 44.0 23.0
Metabletus truncatellus (Linnaeus, 1761) 5 0 2 2 0 3 12 0.0 3.0
Microlestes minutulus (Goeze, 1777) 6 93 34 2 10 5 150 51.5 11.8
Nothiophilus biguttatus (Fabricius, 1779) 0 1 1 8 0 0 10 0.5 2.3
Nothiophilus hypocrite Putzeys, 1866 11 0 16 8 1 3 39 0.5 9.5
Panageus cruxmajor (Linnaeus, 1758) 2 0 0 0 0 1 3 0.0 0.8
Poecilus cupreus* (Linnaeus, 1758) 116 1001 249 19 599 154 2138 800.0 134.5
Pseudoophonus rufipes* (De Geer, 1774) 189 54 104 58 53 74 532 53.5 106.3
Pterostichus caerulescens* (Linnaeus, 1758) 16 66 52 9 230 63 436 148.0 35.0
Pterostichus melanarius* (Illiger, 1798) 474 609 700 779 440 682 3684 524.5 658.8
Pterostichus niger* (Schaller, 1783) 22 13 31 113 1 72 252 7.0 59.5
Pterostichus oblongopunctatus* (Fabricius, 1787) 33 0 0 34 1 1 69 0.5 17.0
Pterostichus strenuus* (Panzer, 1796) 13 0 1 16 0 5 35 0.0 8.8
Pterostichus vernalis* (Panzer, 1796) 0 0 0 0 0 1 1 0.0 0.3
Pterostichus sp. Bonelli, 1810 4 0 0 0 1 0 5 0.5 1.0
Stomis pumicatus (Panzer, 1796) 2 0 0 0 0 0 2 0.0 0.5
Synuchus vivalis* (Illiger, 1798) 8 6 9 77 9 19 128 7.5 28.3
Trechoblemus micros (Herbst, 1784) 0 0 0 0 1 0 1 0.5 0.0
Trechus quadristriatus (Schrank, 1781) 0 5 0 1 17 4 27 11.0 1.3
Trichocellus placidus (Gyllenhal, 1827) 0 0 0 1 0 0 1 0.0 0.3
Sum 3856 3100 3309 3025 2077 1946 17313 2588.5 3034.0
Sum of zoophages 3275 2708 2537 2662 1953 1624 14759 2330.5 2524.5
  1. Abbreviations to Table 1: L – Forest, K – Bushes, R1 – Ditch no. 1, R2 – Ditch no. 2, PI – the first Field, PII – the second Field (for details see “Study area”)
  2. *zoophagous species