# Global Dynamics of a Competitive System of Rational Difference Equations in the Plane

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## Abstract

We investigate global dynamics of the following systems of difference equations Open image in new window , Open image in new window , Open image in new window , where the parameters Open image in new window , Open image in new window , Open image in new window , Open image in new window , and Open image in new window are positive numbers and initial conditions Open image in new window and Open image in new window are arbitrary nonnegative numbers such that Open image in new window . We show that this system has rich dynamics which depend on the part of parametric space. We show that the basins of attractions of different locally asymptotically stable equilibrium points are separated by the global stable manifolds of either saddle points or of nonhyperbolic equilibrium points.

### Keywords

Equilibrium Point Saddle Point Jacobian Matrix Unstable Manifold Global Behavior## 1. Introduction and Preliminaries

In this paper, we study the global dynamics of the following rational system of difference equations:

where the parameters Open image in new window and Open image in new window are positive numbers and initial conditions Open image in new window and Open image in new window are arbitrary numbers. System (1.1) was mentioned in [1] as a part of Open Problem 3 which asked for a description of global dynamics of three specific competitive systems. According to the labeling in [1], system (1.1) is called Open image in new window . In this paper, we provide the precise description of global dynamics of system (1.1). We show that system (1.1) has a variety of dynamics that depend on the value of parameters. We show that system (1.1) may have between zero and two equilibrium points, which may have different local character. If system (1.1) has one equilibrium point, then this point is either locally saddle point or non-hyperbolic. If system (1.1) has two equilibrium points, then the pair of points is the pair of a saddle point and a sink. The major problem is determining the basins of attraction of different equilibrium points. System (1.1) gives an example of semistable non-hyperbolic equilibrium point. The typical results are Theorems 4.1 and 4.5 below.

System (1.1) is a competitive system, and our results are based on recent results developed for competitive systems in the plane; see [2, 3]. In the next section, we present some general results about competitive systems in the plane. The third section deals with some basic facts such as the non-existence of period-two solution of system (1.1). The fourth section analyzes local stability which is fairly complicated for this system. Finally, the fifth section gives global dynamics for all values of parameters.

Let Open image in new window and Open image in new window be intervals of real numbers. Consider a first-order system of difference equations of the form

where Open image in new window

When the function Open image in new window is increasing in Open image in new window and decreasing in Open image in new window and the function Open image in new window is decreasing in Open image in new window and increasing in Open image in new window , the system (1.2) is called *competitive*. When the function Open image in new window is increasing in Open image in new window and increasing in Open image in new window and the function Open image in new window is increasing in Open image in new window and increasing in Open image in new window the system (1.2) is called *cooperative*. A map Open image in new window that corresponds to the system (1.2) is defined as Open image in new window . Competitive and cooperative maps, which are called monotone maps, are defined similarly. *Strongly competitive* systems of difference equations or maps are those for which the functions Open image in new window and Open image in new window are coordinate-wise strictly monotone.

If Open image in new window , we denote with Open image in new window , the four quadrants in Open image in new window relative to Open image in new window , that is, Open image in new window , and so on. Define the *South-East* partial order Open image in new window on Open image in new window by Open image in new window if and only if Open image in new window and Open image in new window . Similarly, we define the *North-East* partial order Open image in new window on Open image in new window by Open image in new window if and only if Open image in new window and Open image in new window . For Open image in new window and Open image in new window , define the *distance from* Open image in new window *to* Open image in new window as Open image in new window . By Open image in new window we denote the interior of a set Open image in new window .

It is easy to show that a map Open image in new window is competitive if it is nondecreasing with respect to the South-East partial order, that is if the following holds:

Competitive systems were studied by many authors; see [4, 5, 6, 7, 8, 9, 10, 11, 12, 13, 14, 15, 16, 17, 18, 19], and others. All known results, with the exception of [4, 6, 10], deal with hyperbolic dynamics. The results presented here are results that hold in both the hyperbolic and the non-hyperbolic cases.

We now state three results for competitive maps in the plane. The following definition is from [18].

Definition 1.1.

Let Open image in new window be a nonempty subset of Open image in new window . A competitive map Open image in new window is said to satisfy condition ( Open image in new window ) if for every Open image in new window , Open image in new window in Open image in new window , Open image in new window implies Open image in new window , and Open image in new window is said to satisfy condition ( Open image in new window ) if for every Open image in new window , Open image in new window in Open image in new window , Open image in new window implies Open image in new window .

The following theorem was proved by de Mottoni and Schiaffino [20] for the Poincaré map of a periodic competitive Lotka-Volterra system of differential equations. Smith generalized the proof to competitive and cooperative maps [15, 16].

Theorem 1.2.

Let Open image in new window be a nonempty subset of Open image in new window . If Open image in new window is a competitive map for which ( Open image in new window ) holds, then for all Open image in new window , Open image in new window is eventually componentwise monotone. If the orbit of Open image in new window has compact closure, then it converges to a fixed point of Open image in new window . If instead ( Open image in new window ) holds, then for all Open image in new window , Open image in new window is eventually componentwise monotone. If the orbit of Open image in new window has compact closure in Open image in new window , then its omega limit set is either a period-two orbit or a fixed point.

The following result is from [18], with the domain of the map specialized to be the Cartesian product of intervals of real numbers. It gives a sufficient condition for conditions ( Open image in new window ) and ( Open image in new window ).

Theorem 1.3 (Smith [18]).

Let Open image in new window be the Cartesian product of two intervals in Open image in new window . Let Open image in new window be a Open image in new window competitive map. If Open image in new window is injective and Open image in new window for all Open image in new window then Open image in new window satisfies ( Open image in new window ). If Open image in new window is injective and Open image in new window for all Open image in new window then Open image in new window satisfies ( Open image in new window ).

Theorem 1.4.

Let Open image in new window be a monotone map on a closed and bounded rectangular region Open image in new window Suppose that Open image in new window has a unique fixed point Open image in new window in Open image in new window Then Open image in new window is a global attractor of Open image in new window on Open image in new window

The following theorems were proved by Kulenović and Merino [3] for competitive systems in the plane, when one of the eigenvalues of the linearized system at an equilibrium (hyperbolic or non-hyperbolic) is by absolute value smaller than Open image in new window while the other has an arbitrary value. These results are useful for determining basins of attraction of fixed points of competitive maps.

Our first result gives conditions for the existence of a global invariant curve through a fixed point (hyperbolic or not) of a competitive map that is differentiable in a neighborhood of the fixed point, when at least one of two nonzero eigenvalues of the Jacobian matrix of the map at the fixed point has absolute value less than one. A region Open image in new window is *rectangular* if it is the Cartesian product of two intervals in Open image in new window .

Theorem 1.5.

Let Open image in new window be a competitive map on a rectangular region Open image in new window . Let Open image in new window be a fixed point of Open image in new window such that Open image in new window is nonempty (i.e., Open image in new window is not the NW or SE vertex of Open image in new window , and Open image in new window is strongly competitive on Open image in new window . Suppose that the following statements are true.

- (a)
The map Open image in new window has a Open image in new window extension to a neighborhood of Open image in new window .

- (b)
The Jacobian matrix of Open image in new window at Open image in new window has real eigenvalues Open image in new window , Open image in new window such that Open image in new window , where Open image in new window , and the eigenspace Open image in new window associated with Open image in new window is not a coordinate axis.

Then there exists a curve Open image in new window through Open image in new window that is invariant and a subset of the basin of attraction of Open image in new window , such that Open image in new window is tangential to the eigenspace Open image in new window at Open image in new window , and Open image in new window is the graph of a strictly increasing continuous function of the first coordinate on an interval. Any endpoints of Open image in new window in the interior of Open image in new window are either fixed points or minimal period-two points. In the latter case, the set of endpoints of Open image in new window is a minimal period-two orbit of Open image in new window .

Corollary 1.6.

If Open image in new window has no fixed point nor periodic points of minimal period-two in Open image in new window , then the endpoints of Open image in new window belong to Open image in new window .

For maps that are strongly competitive near the fixed point, hypothesis b. of Theorem 1.5 reduces just to Open image in new window . This follows from a change of variables [18] that allows the Perron-Frobenius Theorem to be applied to give that, at any point, the Jacobian matrix of a strongly competitive map has two real and distinct eigenvalues, the larger one in absolute value being positive, and that corresponding eigenvectors may be chosen to point in the direction of the second and first quadrants, respectively. Also, one can show that in such case no associated eigenvector is aligned with a coordinate axis.

The following result gives a description of the global stable and unstable manifolds of a saddle point of a competitive map. The result is the modification of Theorem 1.7 from [12].

Theorem 1.7.

In addition to the hypotheses of Theorem 1.5, suppose that Open image in new window and that the eigenspace Open image in new window associated with Open image in new window is not a coordinate axis. If the curve Open image in new window of Theorem 1.5 has endpoints in Open image in new window , then Open image in new window is the global stable manifold Open image in new window of Open image in new window , and the global unstable manifold Open image in new window is a curve in Open image in new window that is tangential to Open image in new window at Open image in new window and such that it is the graph of a strictly decreasing function of the first coordinate on an interval. Any endpoints of Open image in new window in Open image in new window are fixed points of Open image in new window .

The next result is useful for determining basins of attraction of fixed points of competitive maps.

Theorem 1.8.

such that the following statements are true.

- (i)
Open image in new window is invariant, and Open image in new window as Open image in new window for every Open image in new window .

- (ii)
Open image in new window is invariant, and Open image in new window as Open image in new window for every Open image in new window .

If, in addition, Open image in new window is an interior point of Open image in new window and Open image in new window is Open image in new window and strongly competitive in a neighborhood of Open image in new window , then Open image in new window has no periodic points in the boundary of Open image in new window except for Open image in new window , and the following statements are true.

- (iii)
For every Open image in new window there exists Open image in new window such that Open image in new window for Open image in new window .

- (iv)
For every Open image in new window there exists Open image in new window such that Open image in new window for Open image in new window .

## 2. Some Basic Facts

In this section we give some basic facts about the nonexistence of period-two solutions, local injectivity of map Open image in new window at the equilibrium point and Open image in new window condition.

### 2.1. Equilibrium Points

The equilibrium points Open image in new window of system (1.1) satisfy

First equation of System (2.1) gives

Second equation of System (2.1) gives

Now, using (2.2), we obtain

This implies

which is equivalent to

Solutions of (2.6) are

Now, (2.2) gives

The equilibrium points are:

where Open image in new window are given by the above relations.

Note that

The discriminant of (2.6) is given by

Table 1

No equilibrium | |

No equilibrium | |

No equilibrium |

### 2.2. Condition Open image in new window and Period-Two Solution

In this section we prove three lemmas.

Lemma 2.1.

System (1.1) satisfies either Open image in new window or Open image in new window Consequently, the second iterate of every solution is eventually monotone.

Proof.

Lemma 2.2.

System (1.1) has no minimal period-two solution.

Proof.

We show that this system has no other positive solutions except equilibrium points.

From (2.31), we obtain fixed points. In the sequel, we consider (2.32).

Claim.

Assume Open image in new window Then

- (i)
for all values of parameters, Open image in new window

- (ii)
for all values of parameters, Open image in new window

Proof.

which implies that the inequality (2.41) is true.

Now, the proof of the Lemma 2.2 follows from the Claim .

Lemma 2.3.

Proof.

## 3. Linearized Stability Analysis

The Jacobian matrix of the map Open image in new window has the following form:

The value of the Jacobian matrix of Open image in new window at the equilibrium point is

The determinant of (3.2) is given by

The trace of (3.2) is

The characteristic equation has the form

Theorem 3.1.

Assume that Open image in new window Then there exists a unique positive equilibrium Open image in new window which is a saddle point, and the following statements hold.

- (a)
- (b)
If Open image in new window and Open image in new window then Open image in new window and Open image in new window

- (c)
If Open image in new window and Open image in new window then Open image in new window and Open image in new window

- (d)
If Open image in new window and Open image in new window then Open image in new window and Open image in new window

Proof.

which is true since Open image in new window and Open image in new window

establishing the proof of Theorem 3.1.

Since the map Open image in new window is strongly competitive, the Jacobian matrix (3.2) has two real and distinct eigenvalues, with the larger one in absolute value being positive.

The first equation implies that either both eigenvalues are positive or the smaller one is negative.

where Open image in new window

(a) If Open image in new window then the smaller root is negative, that is, Open image in new window

From the last inequality statements Open image in new window and Open image in new window follow.

We now perform a similar analysis for the other cases in Table 1.

Theorem 3.2.

Then Open image in new window exist. Open image in new window is a saddle point; Open image in new window is a sink. For the eigenvalues of Open image in new window the following holds.

- (a)
- (b)
- (c)

Proof.

Note that if Open image in new window and Open image in new window then Open image in new window and Open image in new window which implies Open image in new window , which is a contradiction.

Now, we prove that Open image in new window is a sink.

which is true. (see Theorem 3.1.)

which is always true since Open image in new window and the left side is always negative, while the right side is always positive.

imply that Open image in new window is a saddle point.

The first equation implies that either both eigenvalues are positive or the smaller one is negative.

which is obvious if Open image in new window . Then inequality (3.41) holds. This confirms Open image in new window The other cases follow from (3.41).

Theorem 3.3.

which is non-hyperbolic. The following holds.

- (a)
- (b)

Proof.

Note that Open image in new window

Note that the denominator of (3.48) is always positive.

establishing the proof of the theorem.

Now, we consider the special case of System (1.1) when Open image in new window

In this case system (1.1) becomes

Equilibrium points are solutions of the following system:

The second equation implies

Now, the first equation implies

The map Open image in new window associated to System (3.55) is given by

The Jacobian matrix of the map Open image in new window has the following form:

The value of the Jacobian matrix of Open image in new window at the equilibrium point is

The determinant of (3.61) is given by

The trace of (3.61) is

Theorem 3.4.

of system (1.1), which is a saddle point. The following statements hold.

- (a)
- (b)

Proof.

We prove that Open image in new window is a saddle point.

Hence Open image in new window is a saddle point.

establishing the proof of theorem.

## 4. Global Behavior

Theorem 4.1.

Then system (1.1) has a unique equilibrium point Open image in new window which is a saddle point. Furthermore, there exists the global stable manifold Open image in new window that separates the positive quadrant so that all orbits below this manifold are asymptotic to Open image in new window and all orbits above this manifold are asymptotic to Open image in new window All orbits that start on Open image in new window are attracted to Open image in new window The global unstable manifold Open image in new window is the graph of a continuous, unbounded, strictly decreasing function.

Proof.

The existence of the global stable manifold Open image in new window with the stated properties follows from Theorems 1.5, 1.7, and 1.8 and Lemmas 2.1 and 2.2.

Theorem 4.2.

Then system (1.1) has two equilibrium points: Open image in new window which is a saddle point and Open image in new window which is a sink. Furthermore, there exists the global stable manifold Open image in new window that separates the positive quadrant so that all orbits below this manifold are asymptotic to Open image in new window and all orbits above this manifold are attracted to equilibrium Open image in new window All orbits that start on Open image in new window are attracted to Open image in new window The global unstable manifold Open image in new window is the graph of a continuous, unbounded, strictly decreasing function with end point Open image in new window

Proof.

The existence of the global stable manifold Open image in new window with the stated properties follows from Theorems 1.5, 1.7, and 1.8 and Lemmas 2.1 and 2.2.

Theorem 4.3.

Then system (1.1) has a unique equilibrium Open image in new window which is non-hyperbolic. The sequences Open image in new window , and Open image in new window are eventually monotonic. Every solution that starts in Open image in new window is asymptotic to Open image in new window and every solution that starts in Open image in new window is asymptotic to the equilibrium Open image in new window Furthermore, there exists the global stable manifold Open image in new window that separates the positive quadrant into three invariant regions, so that all orbits below this manifold are asymptotic to Open image in new window and all orbits that start above this manifold are attracted to the equilibrium Open image in new window All orbits that start on Open image in new window are attracted to Open image in new window

Proof.

The existence of the global stable manifold Open image in new window with the stated properties follows from Theorems 1.5, 1.7, and 1.8 and Lemmas 2.1 and 2.2.

Observe that Open image in new window is actually an arbitrary point on the curve Open image in new window , which represents one of two equilibrium curves for system (1.1).

which always holds since the discriminant of the quadratic polynomial on the left-hand side is zero.

Note that Open image in new window and Open image in new window for Open image in new window

Set Open image in new window Then the sequence Open image in new window is increasing and bounded by Open image in new window coordinate of the equilibrium, and the sequence Open image in new window is decreasing and bounded by Open image in new window coordinate of the equilibrium. This implies that Open image in new window converges to the equilibrium as Open image in new window

Set Open image in new window Then the sequence Open image in new window is increasing, and the sequence Open image in new window is decreasing and bounded by Open image in new window coordinate of equilibrium and has to converge. If Open image in new window converges, then Open image in new window has to converge to the equilibrium, which is impossible. This implies that Open image in new window Since Open image in new window then Open image in new window

Since, Open image in new window is asymptotic to Open image in new window then Open image in new window

Theorem 4.4.

Then system (1.1) has a unique equilibrium Open image in new window which is a saddle point. Furthermore, there exists the global stable manifold Open image in new window that separates the positive quadrant so that all orbits below this manifold are asymptotic to Open image in new window and all orbits above this manifold are asymptotic to Open image in new window All orbits that start on Open image in new window are attracted to Open image in new window The global stable manifold Open image in new window is the graph of a continuous, unbounded, strictly increasing function.

Proof.

Theorem 4.5.

Proof.

If the conditions of this theorem are satisfied, then (2.6) implies that there is no real (if the first condition of this theorem is satisfied) or positive equilibrium points (if the second condition of this theorem is satisfied).

This means that sequence Open image in new window is bounded for Open image in new window

for Open image in new window .

Lemma 2.1 implies that subsequences Open image in new window and Open image in new window are eventually monotone.

Since sequence Open image in new window is bounded, then the subsequences Open image in new window and Open image in new window must converge. If the sequences Open image in new window and Open image in new window would converge to finite numbers, then the solution of (1.1) would converge to the period-two solution, which is impossible by Lemma 2.2. Thus at least one of the subsequences Open image in new window and Open image in new window tends to Open image in new window . Assume that Open image in new window as Open image in new window . In view of third equation of (4.25), Open image in new window and in view of first equation of (4.25), Open image in new window which by fourth equation of (4.25) implies that Open image in new window as Open image in new window .

Now, we prove the case when Open image in new window and Open image in new window

It is obvious that these two curves do not intersect, which means that System (4.26) does not possess an equilibrium point.

which always holds.

Set Open image in new window Then the sequence Open image in new window is increasing and the sequence Open image in new window is decreasing. Since Open image in new window is decreasing and Open image in new window then it has to converge. If Open image in new window converges, then Open image in new window has to converge to the equilibrium, which is impossible. This implies that Open image in new window The second equation of System (4.26) implies that Open image in new window

Similarly, we can prove the case Open image in new window

### References

- 1.Camouzis E, Kulenović MRS, Ladas G, Merino O:
**Rational systems in the plane.***Journal of Difference Equations and Applications*2009,**15**(3):303-323. 10.1080/10236190802125264MATHMathSciNetCrossRefGoogle Scholar - 2.Kulenović MRS, Merino O:
**Global bifurcation for discrete competitive systems in the plane.***Discrete and Continuous Dynamical Systems. Series B*2009,**12**(1):133-149.MATHMathSciNetCrossRefGoogle Scholar - 3.Kulenović MRS, Merino O:
**Invariant manifolds for competitive discrete systems in the plane.**to appear in*International Journal of Bifurcation and Chaos*, http://arxiv.org/abs/0905.1772v1 to appear in International Journal of Bifurcation and Chaos, - 4.Burgić Dž, Kalabušić S, Kulenović MRS:
**Nonhyperbolic dynamics for competitive systems in the plane and global period-doubling bifurcations.***Advances in Dynamical Systems and Applications*2008,**3**(2):229-249.MathSciNetGoogle Scholar - 5.Burgić Dž, Kulenović MRS, Nurkanović M:
**Global dynamics of a rational system of difference equations in the plane.***Communications on Applied Nonlinear Analysis*2008,**15**(1):71-84.MATHMathSciNetGoogle Scholar - 6.Clark D, Kulenović MRS:
**A coupled system of rational difference equations.***Computers & Mathematics with Applications*2002,**43**(6-7):849-867. 10.1016/S0898-1221(01)00326-1MATHMathSciNetCrossRefGoogle Scholar - 7.Clark D, Kulenović MRS, Selgrade JF:
**Global asymptotic behavior of a two-dimensional difference equation modelling competition.***Nonlinear Analysis. Theory, Methods & Applications*2003,**52**(7):1765-1776. 10.1016/S0362-546X(02)00294-8MATHMathSciNetCrossRefGoogle Scholar - 8.Franke JE, Yakubu A-A:
**Mutual exclusion versus coexistence for discrete competitive systems.***Journal of Mathematical Biology*1991,**30**(2):161-168. 10.1007/BF00160333MATHMathSciNetCrossRefGoogle Scholar - 9.Franke JE, Yakubu A-A:
**Geometry of exclusion principles in discrete systems.***Journal of Mathematical Analysis and Applications*1992,**168**(2):385-400. 10.1016/0022-247X(92)90167-CMATHMathSciNetCrossRefGoogle Scholar - 10.Garić-Demirović M, Kulenović MRS, Nurkanović M:
**Global behavior of four competitive rational systems of difference equations in the plane.***Discrete Dynamics in Nature and Society*2009,**2009:**-34.Google Scholar - 11.Hirsch MW, Smith H:
**Monotone dynamical systems.**In*Handbook of Differential Equations: Ordinary Differential Equations. Vol. II*. Elsevier, Amsterdam, The Netherlands; 2005:239-357.Google Scholar - 12.Kulenović MRS, Merino O:
**Competitive-exclusion versus competitive-coexistence for systems in the plane.***Discrete and Continuous Dynamical Systems. Series B*2006,**6**(5):1141-1156.MATHMathSciNetCrossRefGoogle Scholar - 13.Kulenović MRS, Nurkanović M:
**Asymptotic behavior of a system of linear fractional difference equations.***Journal of Inequalities and Applications*2005, (2):127-143.Google Scholar - 14.Kulenović MRS, Nurkanović M:
**Asymptotic behavior of a competitive system of linear fractional difference equations.***Advances in Difference Equations*2006,**2006:**-13.Google Scholar - 15.Smith HL:
**Invariant curves for mappings.***SIAM Journal on Mathematical Analysis*1986,**17**(5):1053-1067. 10.1137/0517075MATHMathSciNetCrossRefGoogle Scholar - 16.Smith HL:
**Periodic competitive differential equations and the discrete dynamics of competitive maps.***Journal of Differential Equations*1986,**64**(2):165-194. 10.1016/0022-0396(86)90086-0MATHMathSciNetCrossRefGoogle Scholar - 17.Smith HL:
**Periodic solutions of periodic competitive and cooperative systems.***SIAM Journal on Mathematical Analysis*1986,**17**(6):1289-1318. 10.1137/0517091MATHMathSciNetCrossRefGoogle Scholar - 18.Smith HL:
**Planar competitive and cooperative difference equations.***Journal of Difference Equations and Applications*1998,**3**(5-6):335-357. 10.1080/10236199708808108MATHMathSciNetCrossRefGoogle Scholar - 19.Smith HL:
**Non-monotone systems decomposable into monotone systems with negative feedback.***Journal of Differential Equations*2006,**53:**747-758.MATHGoogle Scholar - 20.de Mottoni P, Schiaffino A:
**Competition systems with periodic coefficients: a geometric approach.***Journal of Mathematical Biology*1981,**11**(3):319-335. 10.1007/BF00276900MATHMathSciNetCrossRefGoogle Scholar

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