Abstract
A new species of the genus Leptoropha (Kotlassiidae, Seymouriamorpha), L. minima sp. nov, is described based on isolated teeth discovered in the Upper Urzhumian beds of the boundary stratotype of the Severodvinian Stage (Republic of Tatarstan, Russia). A sample from the type locality (Monastyrskii Ovrag-D) enabled the tracing of the modification of maxillar and mandibular tooth shape in individuals with skull lengths of 30–60 mm, from the tricuspid pattern to complex polycuspid fan-shaped crowns with a large (up to 14) number of complementary cusps that definitively formed the marginal portions of the protocingulum in the anteriormost jaw teeth. In addition to the type locality, individual teeth attributed to L. minima were identified from the Upper Urzhumian of the Southern Cis-Ural Region (Orenburg Region, Yashkino locality) and the lowermost Severodvinian beds of the Vyatka River basin (Kirov Region, Povoiska locality); findings of teeth of similar morphology at the Sundyr-1 locality (Mari El Republic, Russia) extend the stratigraphic distribution of Leptorophinae to the Upper Severodvinian (Lower Putyatinian) and indicate the presence of the group in the last fauna of the Dinocephalian Tetrapod Superassemblage of Eastern Europe, the Sundyr’ Tetrapod Assemblage.
This is a preview of subscription content, log in via an institution to check access.
Similar content being viewed by others
REFERENCES
Bulanov, V.V., New data on procolophons from the Permian of Eastern Europe, Paleontol. J., 2002, vol. 36, no. 5, pp. 525–530.
Bulanov, V.V., Evolution and systematics of seymouriamorph parareptiles, Paleontol. J., 2003, vol. 37, suppl. no. 1, pp. 1–105.
Bulanov, V.V., Trophic adaptations of seymouriamorphs (Parareptilia) and position of the group in the structure of aquatic communities of the terminal Paleozoic, in Evolyutsiya biosfery i bioraznoobraziya (Evolution of the Biosphere and Biodiversity), Rozhnov, S.V., Ed., Moscow: KMK, 2006, pp. 394–415.
Bulanov, V.V., New finds of tetrapods in the limiting stratotype section of the Severodvinian (Upper Permian) of Eastern Europe near the village of Monastyrskoe (Tatarstan), in Paleontologiya i stratigrafiya permi i triasa Severnoy Yevrazii. Mater. V mezhdunar. konf., posvyashchennoy 150-letiyu so dnya rozhdeniya V. P. Amalitskogo (1860-1917) (Permian and Triassic Paleontology and Stratigraphy of North Eurasia. Mater. V int. Conf. dedicated to the 150th anniversary of V.P. Amalitzky (1860–1917)), Moscow: Paleontol. Inst. Ross. Akad. Nauk, 2010, pp. 53–56.
Bulanov, V.V., New finds of Microphon exiguus (Seymouriamorpha, Kotlassiidae) in the Severodvinian beds of the Sukhona River basin, Russia, Paleontol. J., 2014, vol. 48, no. 6, pp. 633–644.
Golubev, V.K., and Bulanov, V.V., Amphibians of the Permian Sundyr tetrapod assemblage of Eastern Europe, Paleontol. J., 2018, vol. 52, no 6, pp. 639–652.
Golubev, V.K., Kurkin, A.A., and Sennikov, A.G., On the age of the Sundyr Faunal Assemblage of Permian tetrapods of the East European Placket, Uch. Zap. Kazan.Univ. Ser. Est. Nauk., 2015, vol. 157, no. 1, pp. 49–58.
Ivakhnenko, M.F., Permian parareptiles of the USSR, Tr. Paleontol. Inst. Akad. Nauk SSSR, 1987, vol. 223, pp. 1–160.
Minikh, M.G., Minikh, A.V., Molostovskaya, I.I., and Andrushkevich, S.O., On the stratotype section and point of the Severodvinian Stage, Nedra Povolzh’ya i Prikaspiya, 2009, no. 58, pp. 31–38.
Mouraviev, F.A., Arefiev, M.P., Silantiev, V.V., Balaba-nov, Yu.P., Bulanov, V.V., et al., Monastery Ravine section. Stratotype of the Urzhumian and limitotype of the Severodvinian stage, in Type and Reference Sections of the Middle and Upper Permian of the Volga and Kama River Regions: Field Guidebook of XVIII International Congress on the Carboniferous and Permian, Kazan: Kazan. Gos. Univ., 2015, pp. 120–141.
Säilä, L.K., Alpha taxonomy of the Russian Permian procolophonoid reptiles, Acta Palaeontol. Pol., 2009, vol. 54, no. 4, pp. 599–608.
Silantiev, V.V., Arefiev, M.P., Nurgalieva, F.A., Mouraviev, F.A., Bulanov, V.V., et al., Cheremushka section: Parastratotype of the Urzhumian Stage, in Type and Reference Sections of the Middle and Upper Permian of the Volga and Kama River Regions: Field Guidebook of XVIII International Congress on the Carboniferous and Permian, Kazan: Kazan. Gos. Univ., 2015, pp. 70–119.
Tchudinov, P.K., Cotylosaurs from the Shikhovo-Chirki locality, Dokl. Akad. Nauk SSSR, 1955, vol. 103, no. 5, pp. 913–916.
Watson, D.M.S., A sketch classification of the pre-Jurassic tetrapod vertebrates, Proc. Zool. Soc. London, 1917, pp. 167–186.
ACKNOWLEDGMENTS
The author is grateful to A.V. Minikh (Saratov State University) for the materials on Leptorophinae from localities in the Orenburg Region, and assistance in organizing excavations at tetrapod localities in the Monastery Ravine (Monastyrskii Ovrag).
Funding
This work was financially supported by grants nos. 17-04-01937 and 17-04-00410 from the Russian Foundation for Basic Research, the fundamental research program no. 22 “Evolution of the organic world and planetary processes” of the Presidium of the Russian Academy of Sciences, and also from a subsidy allocated within the Russian Government Program of Competitive Growth of Kazan Federal University.
Author information
Authors and Affiliations
Corresponding author
Additional information
Translated by S. Semenova
Rights and permissions
About this article
Cite this article
Bulanov, V.V. A New Leptoropha (Kotlassiidae, Seymouriamorpha) Species from the Upper Urzhumian of European Russia. Paleontol. J. 54, 290–296 (2020). https://doi.org/10.1134/S0031030120030053
Received:
Revised:
Accepted:
Published:
Issue Date:
DOI: https://doi.org/10.1134/S0031030120030053