Abstract
Considerable research has been done on the Ordovician marine fossils from South China, including macrofossils such as brachiopods, graptolites, bivalves, trilobites, some microfossils like conodonts and acritarches. However, radiolarians and ostracods that are also important constituents of the Ordovician marine ecosystem have been paid little attention in this region. In this study, ten radiolarians species belonging to four genera and sixteen ostracods species grouped into nine genera were found from the Hulo Formation at the Hengdu Section of the Jiangshan District, western Zhejiang Province, South China. The fossil-bearing strata belong to the graptolite Pterograptus elegans Zone which indicates the Late Darriwilian Age. This radiolarian fauna is the first record of the Middle Ordovician radiolarian body fossils and also the earliest Ordovician radiolarian fauna reported from South China. The occurrence of Beothuka in this fauna extends the stratigraphic range of the genus to the Upper Darriwilian. Reviews of previous literatures suggest that the diversity of Beothuka was greater during the Early Ordovician, and then declined gradually from the Early Ordovician to the Middle Ordovician before its extinction. The co-occurring ostracod fauna belongs to a shallow-water ecotype. This is contrary to the host lithofacies of the ostracod fauna which represent a deep-water environment. Therefore, these ostacods may have been transported from the shallow-water environment, most likely from the Yangtze carbonate platform. If this conjecture is the truth, then the Jiangshan District was near the shallow-water carbonate platform and received its sediments during the Darriwilian. It is still essential to do more work in the future to better understand the ecology of the Ordovician ostracod fanuas and their role in the sedimentary system of South China.
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References Cited
Aitchison, J. C., Flood, P. G., Malpas, J., 1998. Lowermost Ordovician (Basal Tremadoc) Radiolarians from the Little Port Complex, Western Newfoundland. Geological Magazine, 135(3): 413–419. https://doi.org/10.1017/s001675689800867x
Bandel, K., Becker, G., 1975. Ostracoden aus Paläozoischen–Pelagischen Kalken der Karnischen Alpen (Silurium bis Unterkarbon). Senkenbergiana Lethaea, 56(1): 1–83 (in German)
Berdan, J. M., 1976. Middle Ordovician Leperditicopid Ostracodes from the Ibex Area, Millard County, Western Utah. Brigham Young University College Studies, 23: 37–65
Berdan, J. M., 1984. Leperditicopid Ostracodes from Ordovician Rocks of Kentucky and Nearby States and Characteristic Features of the Order Leperditicopida. United States Government Printing Office, Washington DC. 1–40
Berdan, J. M., 1988. Middle Ordovician (Whiterockian) Palaeocopid and Podocopid Ostracodes from the Ibex Area, Millard County, Western Utah. New Mexico Bureau of Mines and Mineral Resources Memoir, 44: 273–301
Braddy, S. J., Tollerton, V. P., Racheboeuf, P. R., et al., 2004. Eurypterids, Phyllocarids, and Ostracodes. In: Webby, B. D., Paris, F., Droser, M. L., et al., eds., The Great Ordovician Biodiversification Event. Columbia University Press, New York. 255–256. https://doi.org/10.7312/webb12678-026
Buckman, S., Aitchison, J. C., 2001. Middle Ordovician (Llandeilan) Radiolarians from West Junggar, Xinjiang, China. Micropaleontology, 47(4): 359–367. https://doi.org/10.2113/47.4.359
Cui, Z. L., Huan, H., Song, Q. Y., 2000. The Late Ordovician Radiolarian Assemblage of the North Qinling Back–Arc Basin, China. Acta Geologica Sinica, 74(3): 254–258 (in Chinese with English Abstract)
Danelian, T., 1999. Taxonomic Study of Ordovican (Llanvirn–Caradoc) Radiolaria from the Southern Uplands (Scotland, U.K.). Geodiversitas, 21(4): 625–635
Danelian, T., Clarkson, E. N. K., 1998. Ordovician Radiolaria from Bedded Cherts of the Southern Uplands. Scottish Journal of Geology, 34(2): 133–137. https://doi.org/10.1144/sjg34020133
Danelian, T., Floyd, J. D., 2001. Progress in Describing Ordovician Siliceous Biodiversity from the Southern Uplands (Scotland, U.K.). Transactions of the Royal Society of Edinburgh: Earth Sciences, 91(3/4): 489–498. https://doi.org/10.1017/s0263593300008336
Danelian, T., Noble, P., Pouille, L., et al., 2013. Palaeogeographical Distribution of Ordovician Radiolarian Occurrences: Patterns, Significance and Limitations. Geological Society, London, Memoirs, 38(1): 407–413. https://doi.org/10.1144/m38.25
Danelian, T., Popov, L. E., 2003. Ordovician Radiolarian Biodiversity: Insights Based on New and Revised Data from Kazakhstan. Bulletin de la Société Géologique de France, 174(4): 325–335. https://doi.org/10.2113/174.4.325
Danelian, T., Popov, L. E., Tolmacheva, T. Y., et al., 2011. Ordovician Radiolaria and Conodonts from the Peri–Gondwanan Karatau–Naryn Microplate (Sarydzhaz, Eastern Kyrgyzstan). Geobios, 44(6): 587–599. https://doi.org/10.1016/j.geobios.2011.02.006
Fang, Z. J., 2006. An Introduction to Ordovician Bivalves of Southern China, with a Discussion of the Early Evolution of the Bivalvia. Geological Journal, 41(3/4): 303–328. https://doi.org/10.1002/gj.1048
Feng, Z. Z., Peng, Y. M., Jin, Z. K., et al., 2001. Lithofacies Palaeogeography of the Middle and Late Ordovician in South China. Journal of Palaeogeography, 3(4): 10–24 (in Chinese with English Abstract).
Feng, Z. Z., Peng, Y. M., Jin, Z. K., et al., 2003. Lithofacies Palaeogeography of the Middle Ordovician China. Journal of Palaeogeography, 5(3): 263–278 (in Chinese with English Abstract)
Frank, M. S., 1969. Ordovician Ostacode Aparchites Whiteavesi Jones (1889), and Problems of Relationships. Journal of Paleontology, 43(5): 1237–1244
Ghobadi Pour, M., Williams, M., Vannier, J., et al., 2006. Ordovician Ostracods from East Central Iran. Acta Palaeontologica Polonica, 51(3): 551–560
Ghobadi Pour, M., Mohibullah, M., Williams, M., et al., 2011. New, Early Ostracods from the Ordovician (Tremadocian) of Iran: Systematic, Biogeographical and Palaeoecological Significance. Alcheringa: An Australasian Journal of Palaeontology, 35(4): 517–529. https://doi.org/10.1080/03115518.2011.538909
Goto, H., Ishiga, H., 1991. Study of Late Ordovician Radiolarians from the Lachlan Fold Belt, Southeastern Australia. Geological Reports, Shiname University, 10: 57–62
Goto, H., Umeda, M., Ishiga, H., 1992. Late Ordovician Radiolarians from the Lachlan Fold Belt, Southeastern Australia. Memoirs of the Faculty of Science, Shimane University, 26: 145–170
Guo, F. S., 1994. On Collating and Applying of the Lithostratigraphic Units in Jiangshan Area, Zhejiang. Journal of East China Geological Institute, 17(3): 254–263 (in Chinese with English Abstract)
Guo, F. S., 1998. The Ordovician Sedimentary Facies and Palaeogeography and Their Tectonic Controls in Jiangshan, Zhejiang. Sedimentary and Palaeogeography, 18(4): 57–62 (in Chinese with English Abstract)
Henningsmoen, G., 1953. Classification of Paleozoic Straight–Hinge Ostracods. Norske Geologische Tidsskrift, 31: 185–288
Hessland, I., 1949. Lower Ordovician Ostracodes of the Siljan District, Sweden. Bulletin of the Geological Institution of the University of Uppsala, 33: 408
Hou, Y. T., 1956a. Some New Species of Ostracods from Middle Ordovician. Acta Palaeontologica Sinica, 4(3): 355–360 (in Chinese with English Abstract)
Hou, Y. T., 1956b. Ordovician Ostracoda from Western Zhejiang. Acta Palaeontologica Sinica, 4(4): 535–597 (in Chinese with English Abstract)
Hu, K. M., Tang, Z. C., Meng, X. S., et al., 2016. Chronology of Petrogenesis and Mineralization of Datongkeng Porphyry W–Mo Deposit in West Zhejiang. Earth Science, 41(9): 1435–1450. https://doi.org/103799/dqkx2016502 (in Chinese with English Abstract)
Iwata, K., Schmidt, B. L., Leitch, E. C., et al., 1995. Ordovidan Microfossils from the Ballast Formation (Girilambone Group) of New South Wales. Australian Journal of Earth Sciences, 42(4): 371–376. https://doi.org/10.1080/08120099508728208
Jannusson, V., 1957. Middle Ordovician Ostracodes of Central and Southern Sweden. Bulletin Geological Institution of the University of Uppsala, 37: 173–442
Kempf, E. K., 2009. Paraparchites Eskridgensis, New Name for the Ostracode Paraparchites Punctatus Watabe and Kaesler, 2004. Journal of Paleontology, 83(3): 501. https://doi.org/10.1666/09-005.1
Kesling, R. V., Crafts, F. S., Darby, D. G., et al., 1960. Middle Ordovician Black River Ostracods from Michigan, Introduction and Part I. Museum of Paleontology the University of Michigan, XV(13): 293–314
Kesling, R. V., 1960. Middle Ordovician Black River Ostracods from Michigan, Part II. Museum of Paleontology the University of Michigan, XV(15): 349–363
Knüpfer, J., 1968. Ostracoden aus dem Oberen Ordovizium Thüringens. Freiberger Forschung Hefte, C234: 5–29 (in German)
Kozur, H., 1972. Die Bedeutung Triassischer Ostracoden für Stratigraphische und Paläökologische Untersuchungen. Mitteiungen Gesellschaft Geologie Bergbaustelle, 21: 623–660 (in German)
Kuwano, Y., 1987. Early Devonian Conodonts and Ostracodes from Central Japan. Bulletin of Natural. Science Mususem., Tokyo, 13(2): 77–105
Lai, C. G., Jin, R. G., Lin, B. Y., et al., 1993. Biofacies, Sedimentary Facies and Palaeogeographic Characteristics of the Ordovician in the Lower Yangtze Area. Geological Publishing Press, Beijing. 1–84 (in Chinese)
Landing, E., Mohibullah, M., Williams, M., 2013. First Middle Ordovician Ostracods from Western Avalonia: Paleogeographical and Paleoenvironmental Significance. Journal of Paleontology, 87(2): 269–276. https://doi.org/10.1666/12-065r1.1
Levinson, S. A., 1961. New Genera and Species of Bromide (Middle Ordovician) Ostracodes of Oklahoma. Micropaleontology, 7(3): 359–364. https://doi.org/10.2307/1484369
Li, H., 1995. New Genara and Species of Middle Ordovician Nassellaria and Albaillellaria from Baijingsi, Quilian Mountains, China. Scientia Geologica Sinica, 4(3): 331–346
Li, J., Servais, T., Yan, K., et al., 2007. Microphytoplankton Diversity Curves of the Chinese Ordovician. Bulletin de la Societe Geologique de France, 178(5): 399–409. https://doi.org/10.2113/gssgfbull.178.5.399
Liu, C. G., Li, G. R., Wang, D. W., et al., 2016. Middle–Upper Ordovician (Darriwilian–Early Katian) Positive Carbon Isotope Excursions in the Northern Tarim Basin, Northwest China: Implications for Stratigraphic Correlation and Paleoclimate. Journal of Earth Science, 27(2): 317–328. https://doi.org/10.1007/s12583-016-0696-2
Liu, W., Xu, X. S., Feng, X. T., et al., 2010. Radiolarian Siliceous Rocks and Palaoenvironmental Reconstruction for the Upper Ordovician Wufeng Formation in the Middle–Upper Yangtze Area. Sedimentary Geology and Tethyan Geology, 30(3): 65–70 (in Chinese with English Abstract)
Lu, Y., Ma, Y., Wang, Y., et al., 2017. The Sedimentary Response to the Major Geological Events and Lithofacies Characteristics of Wufeng Formation–Longmaxi Formation in the Upper Yangtze Area. Earth Science, 42(7): 1169–1184. https://doi.org/10/3799/dqkx.2017.095 (in Chinese with English Abstract)
Luo, S., Gao, Z., He, Y., et al., 2002. Ordovician Carbonate Contourite Drifts in Hunan and Gansu Provinces, China. In: Stow, D. A. V., ed., Deep–Water Contourite Systems: Modern Drifts and Ancient Series, Seismic and Sedimentary Characteristics. Geological Society Memoirs, London, 22(1): 433–442. https://doi.org/10.1144/gsl.mem.2002.022.01.30
Maletz, J., 2007. The Early Ordovician Beothuka Terranova (Radiolaria) Faunal Assemblage in Western Newfoundland. Paläontologische Zeitschrift, 81(1): 71–82. https://doi.org/10.1007/bf02988380
Maletz, J., 2011. Radiolarian Skeletal Structures and Biostratigraphy in the Early Palaeozoic (Cambrian–Ordovician). Palaeoworld, 20(2/3): 116–133. https://doi.org/10.1016/j.palwor.2010.12.007
Maletz, J., Albanesi, G. L., Voldman, G. G., 2009. Lower Darriwilian Radiolarians from the Argentine Precordillera. Geobios, 42(1): 53–61. https://doi.org/10.1016/j.geobios.2008.09.002
Maletz, J., Bruton, D. L., 2005. The Beothuka Terranova (Radiolaria) Assemblage and Its Importance for the Understanding of Early Ordovician Radiolarian Evolution. Geological Magazine, 142(6): 711–721. https://doi.org/10.1017/s0016756805001391
Maletz, J., Bruton, D. L., 2007. Lower Ordovician (Chewtonian to Castlemainian) Radiolarians of Spitsbergen. Journal of Systematic Palaeontology, 5(3): 245–288. https://doi.org/10.1017/s1477201907002039
Maletz, J., Bruton, D. L., 2008. The Middle Ordovician Proventocitum Procerulum, Radiolarian Assemblage of Spitsbergen and Its Biostratigraphic Correlation. Palaeontology, 51(5): 1181–1200. https://doi.org/10.1111/j.1475-4983.2008.00803.x
Meidla, T., 1996a. Late Ordovician Ostracodes of Estonia. Fossilia Baltica, 2: 1–222
Meidla, T., 1996b. Latest Ordovician Ostracodes of Baltoscandia. Geological Survey of Denmark and Greenland, 20: 65–71
Meidla, T., 2007. Ostracods from the Upper Ordovician Borenshult Fauna, Sweden. GFF, 129(2): 123–132. https://doi.org/10.1080/11035890701292123
Melnikova, L. M., 1999. Ostracodes from the Billingen Horizon (Lower Ordovician) of the Leningrad Region. Paleontological Journal, 33: 147–152
Melnikova, L. M., 2010. Some Ostracodes from the Gur’yanovka Formation (Upper Ordovician) of Northeastern Gorny Altai. Paleontological Journal, 44(4): 399–408. https://doi.org/10.1134/s0031030110040064
Melnikova, L. M., Tolmacheva, T. Y., Ushatinskaya, G. T., 2010. Find of Tremadocian Ostracodes in Cherts of Kazakhstan. Paleontological Journal, 44(1): 36–40. https://doi.org/10.1134/s0031030110010053
Mohibullah, M., Vandenbroucke, T. R. A., Williams, M., et al., 2011. Late Ordovician (Sandbian) Ostracods from the Ardwell Farm Formation, SW Scotland. Scottish Journal of Geology, 47(1): 57–66. https://doi.org/10.1144/0036-9276/01-428
Molostovskaya, I. I., 2010. Ostracodes from the Upper Permian Khivach Formation in Kolyma–Omolon Basin. Paleontological Journal, 44(3): 282–286. https://doi.org/10.1134/s0031030110030068
Nazarov, B. B., Popov, L. Y., 1976. Radiolarians, Inarticulate Brachiopods and Organisms of Uncertain Systematic Positon from the Middle Ordovician of Eastern Kazakhstan. Paleontologicheskiy Zhurnal, 4(4): 33–42
Nazarov, B. B., Popov, L. Y., Apollonov, M. K., 1977. Lower Paleozoic Radiolarians of Kazakhstan. International Geology Review, 19(8): 913–920. https://doi.org/10.1080/00206817709471089
Nazarov, B. B., Popov, L. Y., 1980. Stratigraphy and Fauna of the Siliceous–Carbonate Sequence of the Ordovician of Kazakhstan (Radiolarians and Inarticulate Brachiopods). Trudy Geologicheskiy Institut Akademiy a Nauk SSR, 331: 1–190 (in Russian)
Nazarov, B. B., Ormiston, A. R., 1986. Trends in the Development of Paleozoic Radiolaria. Marine Micropaleontology, 11(1/2/3): 3–32. https://doi.org/10.1016/0377-8398(86)90003-4
Noble, P., Danelian, T., 2004. Ordovician Radiolarian Biodiversity Estimates. In: Webby, B. D., Paris, F., Droser, M. L., et al., eds., The Great Ordovician Biodiversification Event. Columbia University Press, New York. 97–101
Noble, P. J., Webby, B. D., 2009. Katian (Ordovician) Radiolarians from the Malongulli Formation, New South Wales, Australia: A Reexamination. Journal of Paleontology, 83(4): 548–561. https://doi.org/10.1666/08-179r.1
Olempska, E., 1994. Ostracods of the Mójcza Limestone. Palaeontologia Polonica, 53: 129–212
Öpik, A. A, 1935. Ostracoda from the Lower Ordovician Megalaspis–Limestone of Estonia and Russia. Tart Ülikooli j.o. Loodusuurujate Seltsi Aruanded, Annales Societatis Rebus Naturae Investigandis in Universitate Tartu Constitutae, 42: 28–38 (in Russian)
Peng, Y., Peng, Y. B., Lang, X. G., et al., 2016. Marine Carbon–Sulfur Biogeochemical Cycles during the Steptoean Positive Carbon Isotope Excursion (SPICE) in the Jiangnan Basin, South China. Journal of Earth Science, 27(2): 242–254. https://doi.org/10.1007/s12583-016-0694-4
Pouille, L., Danelian, T., Popov, L. E., 2014. A Diverse Upper Darriwilian Radiolarian Assemblage from the Shundy Formation of Kazakhstan: Insights into Late Middle Ordovician Radiolarian Biodiversity. Journal of Micropalaeontology, 33(2): 149–163. https://doi.org/10.1144/jmpaleo2014-008
Reynolds, L., 1987. The Taxonomy and Palaeoecology of Ostracodes from the Devonian Receptaculites Limestone, Taemas, New South Wales, Australia. Palaeontographica A, 162(3–6): 144–203
Salas, M. J., Vannier, J., Williams, M., 2007. Early Ordovician Ostracods from Argentina: Their Bearing on the Origin of Binodicope and Palaeocope Clades. Journal of Paleontology, 81(6): 1384–1395. https://doi.org/10.1666/05-134.1
Salas, M. J., 2011. Early Ordovician (Floian) Ostracods from the Cordillera Oriental, Northwest Argentina. Geological Journal, 46(6): 637–650. https://doi.org/10.1002/gj.1319
Salas, M. J., Vaccari, N. E., 2012. New Insights into the Early Diversification of the Ostracoda: Tremadocian Ostracods from the Cordillera Oriental, Argentina. Acta Palaeontologica Polonica, 57(1): 175–190. https://doi.org/10.4202/app.2009.1110
Schallreuter, R., Suttner, T. J., Hinz–Schallreuter, I., 2005. Late Ordovician Ostracodes from Himalaya and Their Palaeobiogeographic Relations. Berliner Paläobiologische Abhandlungen, 6: 107–108
Schallreuter, R., Hinz–Schallreuter, I., Suttner, T., 2008. New Ordovician Ostracodes from Himalaya and Their Palaeobiological and Palaeogeographical Implications. Revue de Micropaléontologie, 51(3): 191–204. https://doi.org/10.1016/j.revmic.2007.02.003
Schallreuter, R., Hinz–Schalleuter, I., 2009. Ostracod as a Tool for Palaeogeographic Reconstruction in the Ordovician. Institution of Erdwissensch, Berlin. 14
Scotese, C. R., 2001. Atlas of Earth History, Volume 1, Paleogeography, PALEOMAP Project. University of Texas, Arlington. 52
Scott, H. W., 1959. Type Species of Paraparchites Ulrich & Bassler. Journal of Paleontology, 33(4): 679–674
Servais, T., Lehnert, O., Li, J., et al., 2008. The Ordovician Biodiversification: Revolution in the Oceanic Trophic Chain. Lethaia, 41(2): 99–109. https://doi.org/10.1111/j.1502-3931.2008.00115.x
Servais, T., Owen, A. W., Harper, D. A. T., et al., 2010. The Great Ordovician Biodiversification Event (GOBE): The Palaeoecological Dimension. Palaeogeography, Palaeoclimatology, Palaeoecology, 294(3/4): 99–119. https://doi.org/10.1016/j.palaeo.2010.05.031
Shi, C. G., Wang, D. H., 1985. Middle Ordovician Ostracodes from Huanxian, Gansu. Northwest Geoscience, 10: 96–104 (in Chinese with English Abstract)
Siveter, D. J., Rushton, A. W. A., Siveter, D. J., 1995. An Ostracod–Like Arthropod with Appendages Preserved from the Lower Ordovician of England. Lethaia, 28(4): 299–307. https://doi.org/10.1111/j.1502-3931.1995.tb01818.x
Siveter, D. J., Tanaka, G., Farrell, Ú. C., et al., 2014. Exceptionally Preserved 450–Million–Year–Old Ordovician Ostracods with Brood Care. Current Biology, 24(7): 801–806. https://doi.org/10.1016/j.cub.2014.02.040
Song, Y. Y., Zhang, Y. D., Zhang, J., 2013. New Advance in the Late Darriwillian to Early Sandbian Graptolite Biostratigraphy of Western Zhejiang and Northeastern Jiangxi Province, SE China. Journal of Stratigraphy, 37(2): 144–154 (in Chinese with English Abstract)
Spivey, R. C., 1939. Ostracodes from the Maquoketa Shale, Upper Ordovician, of Iowa. Journal of Paleontology, 13(2): 163–175
Sun, Q. Y., 1988. Ordovician Ostracoda from Western Hubei. Acta Micropalaeontologica Sinica, 5(3): 253–266 (in Chinese with English Abstract)
Swain, F. M., 1957. Early Middle Ordovician Ostracoda of the Eastern United States. Part I. Stratigraphic Data and Description of Leperditiidae, Aparchitidae and Leperditellidae. Journal of Paleontology, 31(3): 528–570
Swain, F. M., 1962. Early Middle Ordovician Ostracoda of the Eastern United States. Part II. Leperditellacea, Hollinacea, Kloedenellacea, Bairdiacea and Superfamily Uncertain. Journal of Paleontology, 36(4): 719–744
Tinn, O., Meidla, T., 1999. Ordovician Ostracodes from the Komstad Limestone. Bulletin of the Geological Society of Denmark, 46(1): 25–30
Tinn, O., Meidla, T., 2001. Middle Ordovician Ostracods from the Lanna and Holen Limestones, South–Central Sweden. GFF, 123(3): 129–136. https://doi.org/10.1080/11035890101233129
Tinn, O., Meidla, T., 2003. Ontogeny and Thanatocoenoses of Early Middle Ordovician Palaeocope Ostracode Species Brezelina Palmata (Krause, 1889) and Ogmoopsis Bocki (Öpik, 1935). Journal of Paleontology, 77(1): 64–72. https://doi.org/10.1666/0022-3360(2003)077
Umeda, M., Goto, H., Ishiga, H., 1992. Middle Ordovician Radiolarians from the Lachlan Fold Belt, Southeastern Australia. Memoirs of the Faculty of Science, Shimane University, 26: 131–140
Vannier, J. M. C., Racheboeuf, P. R., Benedetto, J. L., 1995. Silurian–Early Devonian Ostracodes from South America (Argentina, Bolivia): Preliminary Investigations. Journal of Paleontology, 69(4): 752–772. https://doi.org/10.1017/s0022336000035265
Wang, S. Q., 1988. Late Paleozoix Ostracode Association from South China and Their Paleoecological Significance. Acta Palaeontologica Sinica, 27: 91–102 (in Chinese with English Abstract)
Wang, S. Q., 2015. Ordovician and Silurian Ostracoda of China. China University of Technology Press, Hefei. 1–272 (in Chinese)
Wang, X. F., 1989. Palaeogeographic Reconstruction of Ordovician in China and Characteristics of Its Sedimentary Environment and Biofacies. Acta Micropalaeontologica Sinica, 28(2): 234–248 (in Chinese with English Abstract)
Wang, X. F., 2016. Ordovician Tectonic–Paleogeography in South China and Chrono–and Bio–Stratigraphic Division and Correlation. Earth Science Prontiers, 23(6): 253–267 (in Chinese with English Abstract)
Wang, Y. J., 1991. On Progress in the Study of Paleozoic Radiolarians in China. Acta Micropalaeontologica Sinica, 8(3): 237–251 (in Chinese with English Abstract)
Wang, Y. J., 1993. Middle Ordovician Radiolarians from the Pingliang Formation of Gansu Province, China. Micropaleontology Special Publication, 6: 98–114
Wang, Y. J., Cheng, J. F., Zhang, Y. D., 2008. New Radiolarian Genera and Species of Heituao Formation (Ordovician) in the Kuruktag Region, Xinjiang. Acta Palaeontologica Sinica, 47(4): 393–404 (in Chinese with English Abstract)
Wang, Y. J., Zhang, Y. D., 2011. Radiolarian Fauna of the Wufeng Formation (Upper Ordovician) in Lunshan Area, Jiangsu and Its Geological Significance. Acta Micropalaeontologica Sinica, 28(3): 251–260 (in Chinese with English Abstract)
Wang, Z. H., Bergstrom, S. M., Zhang, Y. D., 2015. Upper Ordovician Conodonts from the Yenwashan Formation in the Zhejiang–Jiangxi Border Region, S.E. China and Their Biostratigraphic Significance. Acta Palaeontologica Sinica, 54: 147–157
Williams, M., Stone, P., Siveter, D. J., et al., 2001. Upper Ordovician Ostracods from the Cautley District, Northern England: Baltic and Laurentian Affinities. Geological Magazine, 138(5): 589–607. https://doi.org/10.1017/s0016756801005726
Williams, M., Siveter, D. J., 1996. Lithofacies–Influenced Ostracod Associations in the Middle Ordovician Bromide Formation, Oklahoma, USA. Journal of Micropalaeontology, 15(1): 69–81. https://doi.org/10.1144/jm.15.1.69
Won, M. Z., Iams, W. J., 2011. Earliest Arenig Radiolarians from the Cow Head Group, Western Newfoundland. Journal of Paleontology, 85(1): 156–177. https://doi.org/10.1666/10-102.1
Won, M. Z., Iams, W. J., 2013. Early Ordovician (Early Arenig) Radiolarians from the Cow Head Group and Review of the Little Port Complex Fauna, Western Newfoundland. Palaeoworld, 22(1/2): 10–31. https://doi.org/10.1016/j.palwor.2012.11.001
Won, M. Z., Iams, W. J., 2015a. Early/Middle Arenig (Late Floian) Radiolarian Faunal Assemblages from Cow Head Group, Western Newfoundland. Palaeontographica Abteilung A, 304(1/2/3/4/5/6): 1–63. https://doi.org/10.1127/pala/304/2015/1
Won, M. Z., Iams, W. J., 2015b. Review of the Beothuka Terranova Assemblage and Characteristics of the Middle Arenig (Ordovician, Latest Floian) Radiolarian Assemblage from the Cow Head Group, Newfoundland. Neues Jahrbuch für Geologie und Paläontologie––Abhandlungen, 278(1): 1–21. https://doi.org/10.1127/njgpa/2015/0513
Wu, R. C., Stouge, S., Wang, Z. H., 2012. Conodontophorid Biodiversification during the Ordovician in South China. Lethaia, 45(3): 432–442. https://doi.org/10.1111/j.1502-3931.2011.00303.x
Yan, K., Servais, T., Li, J., et al., 2011. Biodiversity Patterns of Early–Middle Ordovician Marine Microphytoplankton in South China. Palaeogeography, Palaeoclimatology, Palaeoecology, 299(1/2): 318–334. https://doi.org/10.1016/j.palaeo.2010.11.012
Yuan, F. T., Ma, L. X., 1993. Middle Ordovician Ostracodes from the Baota Formation Liangshan, Shanxi. Acta Micropalaeontologica Sinica, 10(2): 223–235 (in Chinese with English Abstract)
Zhan, R. B., Harper, D., 2006. Biotic Diachroneity during the Ordovician Radiation: Evidence from South China. Lethaia, 39(3): 211–226. https://doi.org/10.1080/00241160600799770
Zhan, R. B., Jin, J., 2007. Ordovician–Early Silurian (Llandovery) Stratigraphy and Palaeontology of the Upper Yangtze Platform, South China. Science Press, Beijing. 1–165 (in Chinese with English Abstract)
Zhan, R. B., Jin, J. S., Liu, J. B., 2013. Investigation on the Great Ordovician Biodiversification Event (GOBE): Review and Prospect. Chinese Science Bulletin (Chinese Version), 58(33): 3357–3371. https://doi.org/10.1360/972013-19 (in Chinese)
Zhan, R. B., Rong, J. Y., Cheng, J. H., et al., 2005. Early–Mid Ordovician Brachiopod Diversification in South China. Science in China Series D: Earth Sciences, 48(5): 662–675. https://doi.org/10.1360/03yd0586
Zhan, R. B., Zhang, Y. D., Yuan, W. W., 2007. A New Concept during the Life Process of the Earth––The Great Ordovician Biodiversification Event. Natural Science Process, 17(8): 1006–1014 (in Chinese with English Abstract)
Zhang, Y. D., Chen, X., 2008. The Evolution of Diversity and Environmental Background of the Ordovician Graptolites. Science in China Series D: Earth Sciences, 38(1): 10–21 (in Chinese)
Zhang, Y. D., Chen, X., Goldman, D., 2007. Diversification Patterns of Early and Mid Ordovician Graptolites in South China. Geological Journal, 42(3/4): 315–337. https://doi.org/10.1002/gj.1075
Zhang, Y. D., Zhan, R. B., Fan, J. X., et al., 2009. The Key Scientific Problems of the Research on the Ordovician Radiation. Science in China Series D: Earth Sciences, 39(2): 129–143 (in Chinese)
Zhen, Y. Y., Zhou, Z. Y., 2008. Trilobite Record of China. Science Press, Beijing. 301–330 (in Chinese)
Zheng, N., Song, T. R., Li, T. D., et al., 2012. The Discovery of the Lower Cambrian and Middle Ordovician Radiolaria in the South China Orogenic Belt. Geology in China, 39(1): 260–265 (in Chinese with English Abstract)
Zhou, Z. Y., Yuan, J. L., Zhang, Z. H., et al., 1979. Cambrian Biogeographical Province of Guizhou and the Neighbouring Areas. Journal of Stratigraphy, 4(4): 258–271 (in Chinese)
Zhou, Z. Y., Yuan, W. W., Zhou, Z. Q., 2007. Patterns, Processes and Likely Causes of the Ordovician Trilobite Radiation in South China. Geological Journal, 42(3/4): 297–313. https://doi.org/10.1002/gj.1076
Zhou, Z. Y., Zhen, Y. Y., Zhou, Z. Q., 2008. Division of the Ordovician Geographic Untis of China—A Synopsis. Journal of Palaeogeography, 10(2): 176–182 (in Chinese with English Abstract)
Zong, R. W., Wang, Z. Z., Gong, Y. M., et al., 2015. Ordovician Radiolarians from the Yinisala Ophiolitic Mélange and Their Significance in Western Junggar, Xinjiang, NW China. Science China Earth Sciences, 58(5): 776–783. https://doi.org/10.1007/s11430-014-4974-5
Acknowledgments
This study was supported by the NSFC (No. 41430101) and the Special Fund of the State Key Laboratory of Geological Processes and Mineral Resources, China University of Geosciences in Wuhan (No. MSFGPMR201402). We would like to thank Yi Zhang, Hui Sun and Jiangyan Li for help with the SEM. We are also grateful to Kai Liu and Mingquan Ruan for help with the fieldwork in Zhejiang Province. We really appreciate Fayao Chen for the important suggestions for this paper. Special thanks to the editors and two anonymous reviewers for their constructive comments. The final publication is available at Springer via https://doi.org/10.1007/s12583-017-0951-6..
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Yi, Y., Yuan, A., Aitchison, J.С. et al. Upper Darriwilian (Middle Ordovician) Radiolarians and Ostracods from the Hulo Formation, Zhejiang Provicnce, South China. J. Earth Sci. 29, 886–899 (2018). https://doi.org/10.1007/s12583-017-0951-6
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DOI: https://doi.org/10.1007/s12583-017-0951-6