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Law of the jungle: firm survival and price dynamics in evolutionary markets

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Abstract

In this paper I develop a simple, and general model of supply and demand within which almost any theory of consumer and producer behaviour may be integrated by varying parameters. I then investigate the dynamics of this model and its implications for the theory of market evolution, and show that it unifies a number of insights from evolutionary economics. I extend upon these evolutionary theories and also characterise the distribution of prices across the market and investigate its evolution over time.

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Notes

  1. The unawareness of this book within economics and the lack of a Nobel Prize for Richard Nelson and Sidney Winter is surprising given that a cursory internet search reveals that as of August 2014 the book has received well over 25,000 citations, and is often taken as the foundational document for an entire research program in economics (evolutionary economics) which itself is regularly used as a theoretical basis for business and management research. A similarly cursory search - by way of comparison - reveals as of August 2014 approximately a little under 20,000 citations for George Akerlof’s Nobel Prize winning article which introduced the notion of adverse selection and contributed to the development of the economics of imperfect information.

  2. Insofar as all agents in a standard Walrasian model use exactly the same constrained optimisation decision rule and differ only in their endowments and the preference pre-ordering of their alternatives space.

  3. Those who have sympathies otherwise are directed to the excellent papers of Knudsen (2002; 2004) and Hodgson and Knudsen (2004).

  4. Note that this also opens up a link between an evolutionary model of the market and the notion of unplanned inventories due to effective demand shortfalls which lies at the heart of the macroeconomic analysis of Keynes (1936).

  5. This extension of the model would allow, amongst other things, for the analysis of the role within the market of organisational slack and inventory management, and the definition of firm existence by the factors of production rather than the size of the firm in the market.

  6. See the Oxford price studies around Hall and Hitch (1939); Andrews (1949, 1950, 1964) surveyed in Earl (1995, Ch.8 and 9) and the later studies of Blinder et al. (1998) in which it is axiomatic that firms set prices, and that they set prices according to a variety of different rules.

  7. For instance, if \(h\left (\cdot \right )=\frac {\cdot }{\bar {q}}\) we are talking about an averaging of costs, if h(⋅)=⋅ then we are talking about total costs, and if \(h\left (\cdot \right )=\frac {\partial \cdot }{\partial \bar {q}_{i}}\) we are talking about marginal costs.

  8. Downie rarely uses the word “evolution” in The Competitive Process (1958), much less “Darwinism”, and his focus is more on the production process which generates the capacity for sales of firms, though as Nightingale (1997) has also noted, he does note the importance of past profits to expanding capacity for sales and the ability to lower costs of production.

  9. This view would actually constitute a generalisation of auction models in neoclassical microeconomic theory such as the seminal Myerson (1981) and Myerson and Satterthwaite (1983) mechanism design problems which solve for optimal pricing schemes by assuming the seller knows the distribution of types. The present model requires no such assumption for the analysis of market outcomes.

  10. A recent paper, Richter and Rubenstein (2015) going back to the basics of equilibrium serves to show that equilibrium in psychological sciences must involve much greater assumptions upon the psychology (in economics, the preferences) underlying exchange.

  11. Assuming differentiability, (though this can be relaxed) the elasticity of demand for firm i with respect to the price set by firm j is \(\varepsilon _{d_{i}}^{p_{j}}=\frac {p_{j}}{{q_{i}^{d}}}\frac {\partial {q_{i}^{d}}}{\partial p_{j}}\).

  12. Though with physical interpretations of the variables rather than economic.

  13. “For unto every one that hath shall be given, and he shall have abundance: but from him that hath not shall be taken even that which he hath” - Matthew 25:29, King James Version.

  14. This expression is in fact robust to the exact type of average used, for instance, it does not matter whether \(\bar {q}\) is an arithmetic or geometric mean, or a weighted arithmetic or a weighted geometric mean.

  15. I thank Ulrich Witt for reminding me of this property of replicator dynamic models.

  16. We could incorporate any number on assumptions on the firms’ knowledge and behaviour here from the firm knowing the exact functional form of demand as in neoclassical economics, to the bare minimum that the firm can only observe the growth in its own size.

  17. That is, \(\lim _{F\rightarrow \infty }{\sum }_{j\ne i}\left (\frac {\varepsilon _{d_{i}}^{p_{j}}}{\left (-1\right )}\right )\frac {1}{p_{j}}\frac {\partial p_{j}}{\partial t}<\infty \).

  18. For instance, Einstein’s equations of general relativity, generate interesting insights even without applying the particular set of assumptions (such as those made by Oppenheimer to demonstrate the existence of black holes) which yield an analytical solution.

  19. Strictly speaking of course, an individual growth rate \(\frac {\partial q_{i}}{\partial t}\) would converge to the growth rate of \({q_{i}^{d}}\left (N\quad \left \{ 0\right \}_{j=1}^{F}\quad \left \{ \left \{ {\alpha _{j}^{k}}\right \}_{k=1}^{N_{A}}\right \}_{j=1}^{F}\right )\).

  20. Specifically, this coefficient would be equal to \(\varepsilon \left (\frac {F}{1-F}\right )\frac {\partial p}{\partial t}\), and we would have to assume that elasticities for each firms’ demand and price dynamics are identical across the market so that growth equations can be written in the form \(\delta \left (\frac {1}{p_{i}}-\frac {1}{p_{j}}\right )\).

  21. Note that the inclusion of q i as an amplifying effect for price differentials could be taken as a proxy for the “availability” of information about a particular firm for consumers.

  22. A bounded, typically connected and convex subspace - that is, a multidimensional interval of A.

  23. Though ultimately, whether normal profits prevail in reality is an assumption, since “economic” profits are supposed to include “opportunity” costs of courses of action not taken, and these are vague enough conceptually to be quite malleable in estimating economic profit.

  24. Fisher’s notoriously opaque “fundamental theorem” of natural selection - another core principle within evolutionary mathematics - says that in the absence of other confounding factors growth in average fitness is equal to the variance of fitness (Price 1972b).

  25. In the original evolutionary biology interpretation, these coefficients reflect the ability of organisms with a certain trait to reproduce (Price 1970).

References

  • Alchian A (1950) Uncertainty, evolution and economic theory. J Polit Econ 58(3):211–221

    Article  Google Scholar 

  • Andersen E, Holm J (2013) Directional, stabilizing and disruptive selection: An analysis of aspects of economic evolution based on Price’s equation, dRUID Working Paper No. 13-10

  • Andrews P (1949) A reconsideration of the theory of the individual business. Oxf Econ Pap 1(1):54–89

    Google Scholar 

  • Andrews P (1950) Some aspects of competition in retail trade. Oxf Econ Pap 2(2):137–175

    Google Scholar 

  • Andrews P (1964) On competition in economic theory. MacMillan & Co Ltd., New York

    Book  Google Scholar 

  • Barabasi A, Albert R (1999) Emergence of scaling in random networks. Science 286:509–512

    Article  Google Scholar 

  • Bianconi G, Barabasi A (2001) Bose-einstein condensation in complex networks. Phys Rev Lett 86(24):5632–5635

    Article  Google Scholar 

  • Blinder A, Canetti E, Lebow D (1998) Asking about prices: a new approach to understanding price stickiness. Russell Sage Foundation Publications, New York

    Google Scholar 

  • Cyert R, March J (1963) A behavioral theory of the firm. Prentice-Hall, Inc., Englewood Cliffs

    Google Scholar 

  • Dawkins R (1976) The selfish gene. Oxford University Press, Oxford

    Google Scholar 

  • Debreu G (1959) The theory of value cowles foundation monographs. Yale University Press, New Haven

    Google Scholar 

  • Dixit A, Stiglitz J (1977) Monopolistic competition and optimum product diversity. Am Econ Rev 67(3):297–308

    Google Scholar 

  • Dopfer K, Foster J, Potts J (2004) Micro-meso-macro. J Evol Econ 14 (3):263–279

    Article  Google Scholar 

  • Dopfer K, Potts J (2007) The general theory of economic evolution. Routledge, London

    Book  Google Scholar 

  • Downie J (1958) The Competitive Process. Gerald Duckworth & Co. Ltd.

  • Earl P (1995) Microeconomics for business and marketing. Edward Elgar, Cheltenham

    Google Scholar 

  • Foster J (1993) Economics and the self-organisational approach: Alfred Marshall revisited. Econ J 103(419):975–991

    Article  Google Scholar 

  • Foster J (1997) The analytical foundations of evolutionary economics: From biological analogy to economic self-organization. Struct Chang Econ Dyn 8:427–451

    Article  Google Scholar 

  • Foster J (2005) From simplistic to complex systems in economics. Camb J Econ 29:873–892

    Article  Google Scholar 

  • Foster J (2011) Evolutionary macroeconomics: a research agenda. J Evol Econ 21(1):5–28

    Article  Google Scholar 

  • Gabaix X (2009) Power laws in economics and finance. Annual Review of Economics 1:255–293

    Article  Google Scholar 

  • Hall R, Hitch C (1939) Price theory and business behaviour. Oxford Economic Papers os-2 (1), pp 12–45

  • Hayek F (1945) The use of knowledge in society. Am Econ Rev 35:419–530

    Google Scholar 

  • Hayek F (1989) The pretence of knowledge. Am Econ Rev 79(6):3–7

    Google Scholar 

  • Hodgson G (1997) The evolutionary and non-Darwinian economics of Joseph Schumpeter. J Evol Econ 7(2):131–145

    Article  Google Scholar 

  • Hodgson G (2004) The evolution of institutional economics. Routledge, London

    Book  Google Scholar 

  • Hodgson G (2010) Choice, habit and evolution. J Evol Econ 20:1–18

    Article  Google Scholar 

  • Hodgson G, Knudsen T (2004) The firm as an interactor: Firms as vehicles for habit and routines. J Evol Econ 14:281–307

    Article  Google Scholar 

  • Ironmonger D (1972) New commodities and consumer behaviour. Cambridge University Press, Cambridge

    Google Scholar 

  • Joosten R (2006) Walras and Darwin: an odd couple? J Evol Econ 191:561–573

    Article  Google Scholar 

  • Kahneman D, Tversky A (1979) Prospect theory: an analysis of decision under risk. Econometrica 47(2):263–292

    Article  Google Scholar 

  • Keynes J (1936) The general theory of employment interest and money. Harcourt, Orlando

    Google Scholar 

  • Knudsen T (2002) Economic selection theory. J Evol Econ 12:443–470

    Article  Google Scholar 

  • Knudsen T (2004) General selection theory and economic evolution: The price equation and the replicator/interactor distinction. J Econ Methodol 11(2):147–173

    Article  Google Scholar 

  • Kornai J (1971) Anti-Equilibrium. North-Holland, Cape Town

  • Lancaster K (1966) A new approach to consumer theory. J Polit Econ 74 (2):132–157

    Article  Google Scholar 

  • Lavoie M (1996) Mark-up pricing versus normal cost pricing in post-Keynesian models. Review of Political Economy 8(1):57–66

    Article  Google Scholar 

  • Luttmer EGJ (2007) Selection, growth and the size distribution of firms. Q J Econ 122:1103–1068

    Article  Google Scholar 

  • Malerba F, Nelson R, Orsenigo L, Winter S (1999) “history-friendly” models of industry evolution: The computer industry. Industrial and Coporate Change 8(1):3–40

    Article  Google Scholar 

  • Mas-Collel A, Winston M, Green J (1995) Microeconomic theory. Oxford University Press, Oxford

    Google Scholar 

  • Mata J (2008) The new palgrave dictionary of economics, 2nd Edn. Palgrave Macmillan, Ch. Gibrat’s law

    Google Scholar 

  • McFadden D (2013) The new science of pleasure, NBER Working Paper Series No 18687

  • Metcalfe J (1998) Evolutionary economics and creative destruction. Routledge, London

    Book  Google Scholar 

  • Metcalfe J (2007) Alfred marshall’s mecca: Reconciling the theories of value and development. Econ Rec 83(s1):S1–S22

    Article  Google Scholar 

  • Metcalfe J, Foster J, Ramlogan R (2006) Adaptive economic growth. Camb J Econ 30(1):7–32

    Article  Google Scholar 

  • Myerson R (1981) Optimal auction design. Math Oper Res 6(1):58–73

    Article  Google Scholar 

  • Myerson R, Satterthwaite M (1983) Efficient mechanisms for bilateral trading. J Econ Theory 29:205–281

    Article  Google Scholar 

  • Nelson R, Winter S (1982) An evolutionary theory of economic change. Belknap harvard university press, Cambridge

    Google Scholar 

  • Nightingale J (1997) Anticipating Nelson and Winter: Jack Downie’s theory of evolutionary economic change. J Evol Econ 7:147–167

    Article  Google Scholar 

  • Price G (1970) Selection and covariance. Nature 227:520–521

    Article  Google Scholar 

  • Price G (1972a) Extension of covariance selection mathematics. Ann Hum Genet 35:485–490

    Article  Google Scholar 

  • Price G (1972b) Fisher’s ”fundamental theorem” made clear. Ann Hum Genet 36:129–140

    Article  Google Scholar 

  • Qiu-Dong W (1991) The global solution of the n-body problem. Celest Mech Dyn Astron 50:73–88

    Article  Google Scholar 

  • Raffaelli T (2003) Marshall’s evolutionary economics routledge studies in the history of economics. Routledge, London

    Google Scholar 

  • Read L (1958) I, pencil: My family tree as told to Leonard Read. Foundation for Economic Education, New York

    Google Scholar 

  • Richter M, Rubenstein A (2015) Back to fundamentals: Equilibrium in abstract economies. Am Econ Rev 105(8):2570–2594

    Article  Google Scholar 

  • Schumpeter J (1911) Theory of economic development. Transaction Publishers, Piscataway

    Google Scholar 

  • Selten R (1998) Aspiration adaptation theory. J Math Psychol 42:191–214

    Article  Google Scholar 

  • Silverberg G, Verspagen B (2005) The evolutionary foundations of economics. Cambridge university press, cambridge, pp 506–539. ch. Evolutionary theorizing on economic growth

    Book  Google Scholar 

  • Simon H (1955a) A behavioural model of rational choice. Q J Econ 69(1):99–118

    Article  Google Scholar 

  • Simon H (1955b) On a class of skew distribution functions. Biometrika 42(3-4):425–440

    Article  Google Scholar 

  • Veblen T (1898) Why is economics not an evolutionary science? Q J Econ 12(4):373–397

    Article  Google Scholar 

  • von Bertalanffy L (1950) An outline of general system theory. Br J Philos Sci 1(2):134–165

    Article  Google Scholar 

  • Vromen J (2012) Philosophy of economics. Vol. 13 of handbook of the philosophy of science. North-holland, oxford, ch. Ontological Issues in Evolutionary Economics: The Debate Between Generalised Darwinism and the Continuity Hypothesis, pp 737–764

  • Witt U (1986) Firms’ market behavior under imperfect information and economic natural selection. J Econ Behav Organ 7:265–290

    Article  Google Scholar 

  • Witt U (1999) Bioeconomics as economics from a darwinian perspective. J Bioecon 1:19–34

    Article  Google Scholar 

  • Witt U (2008) What is specific about evolutionary economics? J Evol Econ 18 (5):547–575

    Article  Google Scholar 

Download references

Acknowledgments

The author thanks Harry Bloch, Kurt Dopfer, Peter Earl, John Foster, Stan Metcalfe and Ulrich Witt for their comments and discussions on previous drafts of the present work.

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Correspondence to Brendan Markey-Towler.

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Appendices

Appendix A: An intuitive derivation of the Price equation for economics

One problem with applications of the Price equation to evolutionary economics is that the economic intuition behind it can become difficult to follow. Here I provide a re-derivation (based upon Price (1972a)) of the Price equation using only the terminology of economics. The Price equation serves to decompose the growth rate in average fitness across a population, here this means a decomposition of the growth across the population of the average size of firms. We will begin with the change over one “time period”

$$ E_{q\left( t+1\right)}^{t+1}\left( q\right)-E_{q\left( t\right)}^{t}\left( q\right)=\frac{\sum\nolimits_{i=1}^{F}q_{i} \left( t+1\right)q_{i}\left( t+1\right)}{\sum\nolimits_{j=1}^{F}q_{j}\left( t+1\right)} -\frac{\sum\nolimits_{i=1}^{F}q_{i}\left( t\right)q_{i}\left( t\right)}{\sum\nolimits_{j=1}^{F}q_{j}\left( t\right)} $$
(6.1)

it is important to note that the average here is weighted with respect to the {q i (t)} i = 1, much the same as the distribution of prices, which may seem unusual (weighting firm sizes with firm sizes), though it is less unusual once one grasps that the “second”, or “weighting” firm size included in each term of the above expression is being normalised relative to the size of the market - in a sense, but absolutely not exactly being “cancelled out”. Now, we can define what is often called a “selection coefficient” z i for each individual firm within the market Price (1972a, p.486) which reflects the growth rate of the firms within the marketFootnote 25

$$ z_{i}\left( t\right)=\frac{q_{i}\left( t+1\right)}{q_{i}\left( t\right)}=1+\frac{\partial q_{i}\left( t\right)}{q_{i}\left( t\right)} $$
(6.2)

Now, using this selection coefficient to expand the average size of firms within the market at time t+1, and using the fact that q i (t+1) = q i (t)+Δq i (t) we obtain

$$\begin{array}{@{}rcl@{}} E_{q\left( t+1\right)}^{t+1}\left( q\right)&=&\frac{1}{\sum\nolimits_{j=1}^{F}q_{i}\left( t\right)z_{i}\left( t\right)} \sum\limits_{i=1}^{F}q_{i}\left( t\right)z_{i}\left( t\right)q_{i}\left( t+1\right)\\ &=&\frac{1}{\sum\nolimits_{j=1}^{F}q_{i}\left( t\right)z_{i}\left( t\right)}\sum\limits_{i=1}^{F}q_{i}\left( t\right)z_{i}\left( t\right)\left( q_{i}\left( t\right)+{\Delta} q_{i}\left( t\right)\right) \end{array} $$
(6.3)

Expanding this last equality and re-substituting for q(t+1) we get

$$ E_{q\left( t+1\right)}^{t+1}\left( q\right)=\frac{1}{\sum\limits_{j=1}^{F}q_{i}\left( t\right)z_{i}\left( t\right)}\sum\limits_{i=1}^{F}q_{i}\left( t\right)z_{i}\left( t\right)q_{i}\left( t\right)+\frac{1}{{\sum}_{j=1}^{F}q_{i}\left( t+1\right)}\sum\limits_{i=1}^{F}q_{i}\left( t+1\right){\Delta} q_{i}\left( t\right) $$
(6.4)

Now employing some further mathematical tautologies to manipulate the first term of this expression gives us the following

$$\begin{array}{@{}rcl@{}} E_{q\left( t+1\right)}^{t+1}\left( q\right)&=&\frac{1}{\sum\nolimits_{j=1}^{F}q_{i}\left( t\right)z_{i}\left( t\right)} \frac{\sum\nolimits_{j=1}^{F}q_{i}\left( t\right)}{\sum\nolimits_{j=1}^{F}q_{i}\left( t\right)}\sum\limits_{i=1}^{F}q_{i}\left( t\right)\left[z_{i}\left( t\right)-E_{q\left( t\right)}^{t}\left( z\right)\right]\left[q_{i}\left( t\right)-E_{q\left( t\right)}^{t}\left( q\right)\right]\\ &&+\frac{1}{\sum\nolimits_{j=1}^{F}q_{i}\left( t+1\right)}\sum\limits_{i=1}^{F}q_{i}\left( t+1\right){\Delta} q_{i}\left( t\right)\\ &&+\frac{1}{\sum\nolimits_{j=1}^{F}q_{i}\left( t\right)z_{i}\left( t\right)} \frac{\sum\nolimits_{j=1}^{F}q_{i}\left( t\right)}{\sum\nolimits_{j=1}^{F}q_{i}\left( t\right)} \left[\sum\limits_{i=1}^{F}q_{i}\left( t\right)z_{i}\left( t\right)E_{q\left( t\right)}^{t}\left( q\right)\right.\\ &&\qquad\qquad\qquad\qquad\qquad\qquad\qquad\quad\left.+\sum\limits_{i=1}^{F}q_{i}\left( t\right)\left[q_{i}\left( t\right)-E_{q\left( t\right)}^{t}\left( q\right)\right]E_{q\left( t\right)}^{t}\left( z\right)\right]\\ \end{array} $$
(6.5)

Notice that when we expand this last term out carefully it collapses to \(E_{q\left (t\right )}^{t}\left (q\right )\). We can now recognise that this equation is reducible using the quotidian statistical concepts of covariance and expectation

$$ E_{q\left( t\right)}^{t+1}\left( q\right)=\frac{1}{E_{q\left( t\right)}^{t}\left( z\right)}\text{Cov}_{q\left( t\right)}^{t}\left( z\quad q\right)+E_{q\left( t+1\right)}^{t}\left( {\Delta} q\left( t\right)\right)+E_{q\left( t\right)}^{t}\left( q\right) $$
(6.6)

Now subtracting \(E_{q\left (t\right )}^{t}\left (q\right )\) from both sides gives us

$$ E_{q\left( t\right)}^{t+1}\left( q\right)-E_{q\left( t\right)}^{t}\left( q\right)=\frac{1}{E_{q\left( t\right)}^{t}\left( z\right)}\text{Cov}_{q\left( t\right)}^{t}\left( z\quad q\right)+E_{q\left( t+1\right)}^{t}\left( {\Delta} q\left( t\right)\right) $$
(6.7)

Which can be further reduced to the elegant Price equation by generalising to an arbitrary time period and (for the sake of it) assuming differentiability in time

$$ \frac{\partial E_{q}\left( q\right)}{\partial t}=\frac{1}{E_{q}\left( q\right)}\text{Cov}_{q}\left( z\quad q\right)+E_{q}\left( \frac{\partial q}{\partial t}\right) $$
(6.8)

Now, inputting \(z_{i}=1+\frac {1}{q}\frac {\partial q}{\partial t}\partial t\) into the definition of covariance gives us

$$\frac{\partial E_{q}\left( q\right)}{\partial t}=\frac{1}{E_{q}\left( q\right)}\text{Cov}_{q}\left( \frac{\partial t}{q}\frac{\partial q}{\partial t}\quad q\right)+E_{q}\left( \frac{\partial q}{\partial t}\right) $$

Appendix B: Proofs of theorems

1.1 Proof of Theorem 1

Proof

If it is the case that p j = 0 ∀ j then by the definition of demand elasticity \(\varepsilon _{d_{i}}^{p_{j}}=\frac {p_{j}}{{q_{i}^{d}}}\frac {\partial {q_{i}^{d}}}{\partial p_{j}}=0\,\forall \,j\). This implies that \(\left (\frac {\varepsilon _{d_{i}}^{p_{i}}}{F-1}\right )\frac {1}{p_{i}}\frac {\partial p_{i}}{\partial t}=0\) and \(\left (\frac {\varepsilon _{d_{i}}^{p_{j}}}{\left (-1\right )}\right )\frac {1}{p_{j}}\frac {\partial p_{j}}{\partial t}=0\,\forall \,j\ne i\), since by induction \(0\times \lim _{x\rightarrow \infty }x=0\). Inserting this into Eq. 3.3 yields the result. Similarly, we need only input directly the assumption p j / t = 0, ∀ j ∈ {1,…, F} into Eq. 3.3 to obtain the result. □

1.2 Proof of Theorem 2

Proof

First take Eq. 3.3 and input the assumptions of normal and substitutable goods, a zero population growth rate and unchanging product attributes. Then the growth rate of the firm can be expressed as

$$ \frac{\partial q_{i}}{\partial t}={q_{i}^{d}}\sum\limits_{j\ne i}\left[\left( \frac{\varepsilon_{d_{i}}^{p_{i}}}{F-1}\right)\frac{1}{p_{i}}\frac{\partial p_{i}}{\partial t}-\left( \frac{\varepsilon_{d_{i}}^{p_{j}}}{\left( -1\right)}\right)\frac{1}{p_{j}}\frac{\partial p_{j}}{\partial t}\right] $$
(6.9)

Now, it is a mere fact of logic that any output bought must have been sold, so \({q_{i}^{d}}=q_{i}\). But note that \(q_{i}=0\implies \frac {\partial q_{i}}{\partial t}\ne 0\Leftrightarrow {\sum }_{j\ne i}\left [\left (\frac {\varepsilon _{d_{i}}^{p_{i}}}{F-1}\right )\frac {1}{p_{i}}\frac {\partial p_{i}}{\partial t}-\left (\frac {\varepsilon _{d_{i}}^{p_{j}}}{\left (-1\right )}\right )\frac {1}{p_{j}}\frac {\partial p_{j}}{\partial t}\right ]\ne 0\). Since the laws of physics decree that q i ≥0 therefore, it is the case that

$$ \frac{\partial q_{i}}{\partial t}>0\Leftrightarrow\sum\limits_{j\ne i}\left[\left( \frac{\varepsilon_{d_{i}}^{p_{i}}}{F-1}\right)\frac{1}{p_{i}}\frac{\partial p_{i}}{\partial t}-\left( \frac{\varepsilon_{d_{i}}^{p_{j}}}{\left( -1\right)}\right)\frac{1}{p_{j}}\frac{\partial p_{j}}{\partial t}\right]>0 $$
(6.10)

Now suppose that we can partition the set of firms {1,…, F} into two sets B and ¬B such that ∀ jB we have \(\left [\left (\frac {\varepsilon _{d_{i}}^{p_{i}}}{F-1}\right )\frac {1}{p_{i}}\frac {\partial p_{i}}{\partial t}-\left (\frac {\varepsilon _{d_{i}}^{p_{j}}}{\left (-1\right )}\right )\frac {1}{p_{j}}\frac {\partial p_{j}}{\partial t}\right ]>0\). Then condition (6.10) holds if and only if

$$\begin{array}{@{}rcl@{}} \sum\limits_{j\in B}\left[\left( \frac{\varepsilon_{d_{i}}^{p_{i}}}{F-1}\right)\frac{1}{p_{i}}\frac{\partial p_{i}}{\partial t}-\left( \frac{\varepsilon_{d_{i}}^{p_{j}}}{\left( -1\right)}\right)\frac{1}{p_{j}}\frac{\partial p_{j}}{\partial t}\right]>\sum\limits_{j\notin B}\left[\left( \frac{\varepsilon_{d_{i}}^{p_{i}}}{F-1}\right)\frac{1}{p_{i}}\frac{\partial p_{i}}{\partial t}-\left( \frac{\varepsilon_{d_{i}}^{p_{j}}}{\left( -1\right)}\right)\frac{1}{p_{j}}\frac{\partial p_{j}}{\partial t}\right]\\ \end{array} $$
(6.11)

1.3 Proof of Theorem 3

Proof

This condition consists of a simple rearrangement of Eq. 3.3 when restricted to be greater than zero, so

$$ \frac{\partial {q_{i}^{d}}}{\partial N}\frac{\partial N}{\partial t}+{q_{i}^{d}}\sum\limits_{j\ne i}\left[\left( \frac{\varepsilon_{d_{i}}^{p_{i}}}{F-1}\right)\frac{1}{p_{i}}\frac{\partial p_{i}}{\partial t}-\left( \frac{\varepsilon_{d_{i}}^{p_{j}}}{\left( -1\right)}\right)\frac{1}{p_{j}}\frac{\partial p_{j}}{\partial t}\right]+\sum\limits_{j=1}^{F}\sum\limits_{k=1}^{N_{A}}\frac{\partial {q_{i}^{d}}}{{\partial\alpha_{j}^{k}}}\frac{{\partial\alpha_{j}^{k}}}{\partial t}>0 $$
(6.12)

when rearranged gives us

$$\begin{array}{@{}rcl@{}} &&\sum\limits_{k=1}^{N_{A}}\frac{\partial {q_{i}^{d}}}{{\partial\alpha_{i}^{k}}}\frac{{\partial\alpha_{i}^{k}}}{\partial t}>{q_{i}^{d}}\sum\limits_{j\ne i}\left[\left( \frac{\varepsilon_{d_{i}}^{p_{j}}}{\left( -1\right)}\right)\frac{1}{p_{j}}\frac{\partial p_{j}}{\partial t}-\left( \frac{\varepsilon_{d_{i}}^{p_{i}}}{F-1}\right)\frac{1}{p_{i}}\frac{\partial p_{i}}{\partial t}\right]\\ &&-\sum\limits_{j\ne i}^{F}\sum\limits_{k=1}^{N_{A}}\left[\frac{\partial {q_{i}^{d}}}{{\partial\alpha_{j}^{k}}}\frac{{\partial\alpha_{j}^{k}}}{\partial t}\right]-\frac{\partial {q_{i}^{d}}}{\partial N}\frac{\partial N}{\partial t} \end{array} $$
(6.13)

1.4 Proof of Theorem 4

Proof

The market share of firm i with respect to output is given by \(s_{i}=\frac {q_{i}}{{\sum }_{j=1}^{F}q_{j}}\), and its dynamics can be readily obtained by simple application of the quotient rule

$$ \frac{\partial s_{i}}{\partial t}=\frac{\frac{\partial q_{i}}{\partial t}\sum\nolimits_{j=1}^{F}q_{j}-q_{i}\frac{\partial\sum\nolimits_{j=1}^{F}q_{j}}{\partial t}}{\left( \sum\nolimits_{j=1}^{F}q_{j}\right)^{2}} $$
(6.14)

It is fairly straightforward then to confirm that market share only grows if the firm grows at a faster rate than output across the market

$$ \frac{\partial s_{i}}{\partial t}>0\Leftrightarrow\frac{1}{q_{i}}\frac{\partial q_{i}}{\partial t}>\frac{1}{\sum\nolimits_{j=1}^{F}q_{j}}\frac{\partial\sum\nolimits_{j=1}^{F}q_{j}}{\partial t} $$
(6.15)

using the additive law of differential calculus we can expand this into a slightly more tractable (in view of the model above) expression

$$ \frac{\partial q_{i}}{\partial t}>\frac{s_{i}}{1-s_{i}}\sum\limits_{j\ne i}\frac{\partial q_{j}}{\partial t} $$
(6.16)

Now substituting in Eq. 3.3 for the growth rate of firm size we obtain

$$\begin{array}{@{}rcl@{}} &&\frac{\partial {q_{i}^{d}}}{\partial N}\frac{\partial N}{\partial t}+{q_{i}^{d}}\sum\limits_{j\ne i}\left[\left( \frac{\varepsilon_{d_{i}}^{p_{i}}}{F-1}\right)\frac{1}{p_{i}}\frac{\partial p_{i}}{\partial t}-\left( \frac{\varepsilon_{d_{i}}^{p_{j}}}{\left( -1\right)}\right)\frac{1}{p_{j}}\frac{\partial p_{j}}{\partial t}\right]+\sum\limits_{j=1}^{F}\sum\limits_{k=1}^{N_{A}}\frac{\partial {q_{i}^{d}}}{{\partial\alpha_{j}^{k}}}\frac{{\partial\alpha_{j}^{k}}}{\partial t}\\ &&\quad>\frac{s_{i}}{1-s_{i}}\sum\limits_{j\ne i}\left\{ \frac{\partial {q_{j}^{d}}}{\partial N}\frac{\partial N}{\partial t}+{q_{j}^{d}}\sum\limits_{n\ne j}\left[\left( \frac{\varepsilon_{d_{j}}^{p_{j}}}{F-1}\right)\frac{1}{p_{j}}\frac{\partial p_{j}}{\partial t}-\left( \frac{\varepsilon_{d_{j}}^{p_{n}}}{\left( -1\right)}\right)\frac{1}{p_{n}}\frac{\partial p_{n}}{\partial t}\right]\right.\\ &&\quad\left.+\sum\limits_{n=1}^{F}\sum\limits_{k=1}^{N_{A}}\frac{\partial {q_{j}^{d}}}{{\partial\alpha_{n}^{k}}}\frac{{\partial\alpha_{n}^{k}}}{\partial t}\right\} \end{array} $$
(6.17)

We can reduce this inequation by isolating behavioural and firm strategy terms which refer to firm i as follows

$$\begin{array}{@{}rcl@{}} \frac{\partial {q_{i}^{d}}}{\partial N}\frac{\partial N}{\partial t}&+&\frac{s_{i}}{1-s_{i}}\sum\limits_{j\ne i}\frac{\partial {q_{j}^{d}}}{\partial N}\frac{\partial N}{\partial t}\\ &+&{q_{i}^{d}}\sum\limits_{j\ne i}\left[\left( \frac{\varepsilon_{d_{i}}^{p_{i}}}{F-1}\right)\frac{1}{p_{i}}\frac{\partial p_{i}}{\partial t}-\left( \frac{\varepsilon_{d_{i}}^{p_{j}}}{\left( -1\right)}\right)\frac{1}{p_{j}}\frac{\partial p_{j}}{\partial t}\right]\\ &-&\sum\limits_{j\ne i}\frac{s_{i}}{1-s_{i}}{q_{j}^{d}}\left[\left( \frac{\varepsilon_{d_{j}}^{p_{j}}}{F-1}\right)\frac{1}{p_{j}}\frac{\partial p_{j}}{\partial t}-\left( \frac{\varepsilon_{d_{j}}^{p_{i}}}{\left( -1\right)}\right)\frac{1}{p_{i}}\frac{\partial p_{i}}{\partial t}\right]\\ &-&\sum\limits_{j\ne i}\frac{s_{i}}{1-s_{i}}\left\{ {q_{j}^{d}}\sum\limits_{n\ne j,i}\left[\left( \frac{\varepsilon_{d_{j}}^{p_{j}}}{F-1}\right)\frac{1}{p_{j}}\frac{\partial p_{j}}{\partial t}-\left( \frac{\varepsilon_{d_{j}}^{p_{n}}}{\left( -1\right)}\right)\frac{1}{p_{n}}\frac{\partial p_{n}}{\partial t}\right]\right\} \\ &+&\sum\limits_{k=1}^{N_{A}}\frac{\partial {q_{i}^{d}}}{{\partial\alpha_{i}^{k}}}\frac{{\partial\alpha_{i}^{k}}}{\partial t}+\sum\limits_{j\ne i}\sum\limits_{k=1}^{N_{A}}\frac{\partial {q_{i}^{d}}}{{\partial\alpha_{j}^{k}}}\frac{{\partial\alpha_{j}^{k}}}{\partial t}-\frac{s_{i}}{1-s_{i}}\sum\limits_{j\ne i}\sum\limits_{k=1}^{N_{A}}\frac{\partial {q_{j}^{d}}}{{\partial\alpha_{i}^{k}}}\frac{{\partial\alpha_{i}^{k}}}{\partial t}\\ &-&\frac{s_{i}}{1-s_{i}}\sum\limits_{j\ne i}\sum\limits_{n\ne i}\sum\limits_{k=1}^{N_{A}}\frac{\partial {q_{j}^{d}}}{{\partial\alpha_{n}^{k}}}\frac{{\partial\alpha_{n}^{k}}}{\partial t}>0 \end{array} $$
(6.18)

if we multiply and divide every term in \({\sum }_{j\ne i}^{F}{\sum }_{k=1}^{N_{A}}\frac {\partial {q_{i}^{d}}}{{\partial \alpha _{i}^{k}}}\frac {{\partial \alpha _{i}^{k}}}{\partial t}\) by \(\frac {{\partial \alpha _{i}^{k}}}{{\partial \alpha _{i}^{k}}}\) we can by grouping like terms obtain

$$\begin{array}{@{}rcl@{}} &&\frac{\partial N}{\partial t}\left[\frac{\partial {q_{i}^{d}}}{\partial N}-\frac{s_{i}}{1-s_{i}}\sum\limits_{j\ne i}\frac{\partial {q_{j}^{d}}}{\partial N}\right]\\ &&+{q_{i}^{d}}\sum\limits_{j\ne i}\left\{ \frac{1}{p_{i}}\frac{\partial p_{i}}{\partial t}\left[\left( \frac{\varepsilon_{d_{i}}^{p_{i}}}{F-1}\right)+\frac{s_{i}}{1-s_{i}}\frac{{q_{j}^{d}}}{{q_{i}^{d}}}\left( \frac{\varepsilon_{d_{j}}^{p_{i}}}{\left( -1\right)}\right)\right]-\frac{1}{p_{j}}\frac{\partial p_{j}}{\partial t}\left[\left( \frac{\varepsilon_{d_{i}}^{p_{j}}}{\left( -1\right)}\right)-\frac{s_{i}}{1-s_{i}}\frac{{q_{j}^{d}}}{{q_{i}^{d}}}\left( \frac{\varepsilon_{d_{j}}^{p_{j}}}{F-1}\right)\right]\right\} \\ &&-\frac{s_{i}}{1-s_{i}}\sum\limits_{j\ne i}\left\{ {q_{j}^{d}}{\sum}_{n\ne j,i}\left[\left( \frac{\varepsilon_{d_{j}}^{p_{j}}}{F-1}\right)\frac{1}{p_{j}}\frac{\partial p_{j}}{\partial t}-\left( \frac{\varepsilon_{d_{j}}^{p_{n}}}{\left( -1\right)}\right)\frac{1}{p_{n}}\frac{\partial p_{n}}{\partial t}\right]\right\} \\ &&+\sum\limits_{k=1}^{N_{A}}\frac{{\partial\alpha_{i}^{k}}}{\partial t}\left[\frac{\partial {q_{i}^{d}}}{{\partial\alpha_{i}^{k}}}-\frac{s_{i}}{1-s_{i}}\sum\limits_{j\ne i}\left( \sum\limits_{k=1}^{N_{A}}\frac{\partial {q_{j}^{d}}}{{\partial\alpha_{i}^{k}}}-\frac{\partial {q_{i}^{d}}}{{\partial\alpha_{j}^{k}}}\frac{{\partial\alpha_{j}^{k}}}{{\partial\alpha_{i}^{k}}}\right)\right]-\frac{s_{i}}{1-s_{i}}\sum\limits_{j\ne i}\left\{ \sum\limits_{n\ne i}\sum\limits_{k=1}^{N_{A}}\frac{\partial {q_{j}^{d}}}{{\partial\alpha_{n}^{k}}}\frac{{\partial\alpha_{n}^{k}}}{\partial t}\right\} >0\\ \end{array} $$
(6.19)

Which when we rearrange to isolate on the left hand side any market forces over which firm i has influence gives us our result. □

1.5 Proof of Lemma 1

Proof

This proof is rather trivial and well known to mathematicians and statisticians, but will be included for the sake of rigour. Firstly, if the price distribution is degenerate then

$$ E_{q}\left( p\right)=\frac{\sum\nolimits_{i=1}^{F}q_{i}p_{i}}{\sum\nolimits_{j=1}^{F}q_{j}}= \frac{\sum\nolimits_{i\in{\Xi}\left( p\right)}q_{i}p_{i}}{\sum\nolimits_{i\in{\Xi}\left( p\right)}q_{j}} $$
(6.20)

and by the definition of Ξ(p), p i = pi ∈ Ξ(p) and so ∀ i ∈ Ξ(p), E q (p) = p i , while by the definition of degeneracy this must be true for any i:q i > 0. The zero variance of the distribution follows immediately by inputting p i = E q (p) ∀ i:q i > 0 into the definition of Var q (p). Conversely if it is to be the case that p i = E q (p) ∀ i:q i > 0, and Var q (p) = 0 then by the definition of the first and second moments it must be the case that ∃{i}⊂{1,…, F}:p i = E q (p) ∀ i ∈ {i} and for any i not in this set q i = 0, which is the definition of degeneracy. □

1.6 Proof of Theorem 5

Proof

In order for a uniform price to prevail across the market the price distribution must be degenerate by Lemma 1. But for the price distribution to be degenerate it can seen that we will need the dynamics of the market to lead to convergence to a situation in which \({\sum }_{i\in {\Xi }\left (p\right )}s_{i}=1\), since

$$\sum\limits_{i\in{\Xi}\left( E_{q}\left( p\right)\right)}s_{i}=1\iff\sum\limits_{i\in{\Xi}\left( E_{q}\left( p\right)\right)} \frac{q_{i}}{\sum\limits_{j=1}^{F}q_{j}}=\frac{1}{\sum\limits_{j=1}^{F}q_{j}}\sum\limits_{i\in{\Xi}\left( E_{q}\left( p\right)\right)}q_{i}=1\iff{\sum}_{i\in{\Xi}\left( E_{q}\left( p\right)\right)}q_{i}={\sum}_{j=1}^{F}q_{j} $$

But given that Ξ(E q (p))⊂{1,…, F}, for each q i on the left hand side there is a corresponding q i on the right hand side, and so for this to be the case requires that q i = 0 ∀ i∉Ξ(E q (p)), which is the key to the distribution being degenerate by the definition referred to in Lemma 1. So saying that \({\sum }_{i\in {\Xi }\left (p\right )}s_{i}=1\) is identical to saying that the price distribution is degenerate. Hence if it is to be the case that in the market there is convergence to a situation in which \({\sum }_{i\in {\Xi }\left (E_{q}\left (p\right )\right )}s_{i}=1\), it must also be the case by definition of market share that there is a convergence to a situation in which s i = 0 ∀ i∉Ξ(E q (p)). But then for it to be the case that limt s i = 0 ∀ i∉Ξ(E q (p)), for each and every i∉Ξ(E q (p)) from an initial market share of s i we must have

$$s_{i}+{\lim}_{T\rightarrow\infty}{{\int}_{t}^{T}}\frac{\partial s_{i}}{\partial t}\partial t=0 $$
$$\implies s_{i}=-{\lim}_{T\rightarrow\infty}{{\int}_{t}^{T}}\frac{\partial s_{i}}{\partial t}\partial t $$

It should be fairly clear that this will hold if and only if there are a sufficient number of time periods for which \(\frac {\partial s_{i}}{\partial t}<0\) so that \(\lim _{T\rightarrow \infty }{{\int }_{t}^{T}}\frac {\partial s_{i}}{\partial t}\partial t<0\). That is, there must exist a set of time periods \(T_{i\notin {\Xi }\left (p\right )}\subset \mathbb {R}_{+}\) for each and every i∉Ξ(E q (p)) such that

$$\frac{\partial s_{i}}{\partial t}<0\,\forall\,t\in T_{i\notin{\Xi}\left( E_{q}\left( p\right)\right)} $$

that is, for these periods Theorem 4 are violated for each firm i, and

$$s_{i}=-{\lim}_{T\rightarrow\infty}\left\{ {\int}_{t\in T_{i\notin{\Xi}\left( E_{q}\left( p\right)\right)}}\frac{\partial s_{i}}{\partial t}\partial t-{\int}_{t\notin T_{i\notin{\Xi}\left( E_{q}\left( p\right)\right)}}\frac{\partial s_{i}}{\partial t}\partial t\right\} $$

1.7 Proof of Corollary 1

Proof

This is simply the reverse argument of Theorem 5 for the emergence of a dominant firm rather than the non-survival of those which do not conform the uniform price. □

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Markey-Towler, B. Law of the jungle: firm survival and price dynamics in evolutionary markets. J Evol Econ 26, 655–696 (2016). https://doi.org/10.1007/s00191-016-0446-8

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