Paläontologische Zeitschrift

, Volume 77, Issue 2, pp 281–292 | Cite as

Theropods (dinosauria, saurischia) from the middle Jurassic Toutunhe Formation of the Southern Junggar Basin, NW China

  • Michael W. Maisch
  • Andreas T. Matzke


From the Middle Jurassic (?Bathonian-Callovian) Toutunhe Formation of Liuhonggou, SW of Urumqi in the southern Junggar Basin (Xinjiang Uygur Autonomous Region, People’s Republic of China) theropod material is described in detail for the first time. A large carnosaur is represented by five tooth crowns from different parts of the dentition. The size of the teeth indicates an animal 6–8 m in total length. Teeth and postcranial material from the Toutunhe Formation of Sangonghe River, E of Urumqi, belong to the same or a closely related taxon. A second, smaller theropod with an estimated entire length of 3 m is represented by another tooth crown, re-markable for its bifurcated mesial carina (genetically induced abnormality ?). It is not identifiable but shows greatest similarities to members of the Coelophysoidea. The distal end of a fibula could belong to the same taxon. Two new components are thus added to the dinosaur fauna of the Toutunhe Formation, hitherto restricted to a nodosaurid and unidentified (“megalosaurid”) theropod material.


Dinosauria Theropoda Middle Jurassic Junggar Basin Toutnhe Formation 


Aus der mitteljurassischen (?Bathonium-Callovium) Toutunhe-Formation von Liuhonggou, SW Urumqi, im südlichen Junggar-Becken (Autonome Region Xinjiang [ehem. Sinkiang], NW China) wird erstmals detailliert Material von carnivoren Dinosauriern (Theropoda) beschrieben. Ein großwüchsiger Carnosaurier ist durch fünf Zahnkronen aus verschiedenen Bereichen des Gebisses repräsentiert. Die Größe der Zähne lässt auf ein Tier von 6–8 m Gesamtlänge schließen. Zähne und Poscranialmaterial aus der Toutunhe-Formation des Sangonghe River, NE Urumqi, gehören zum gleichen oder einem nahe verwandten Taxon. Ein zweiter, kleinerer Theropode mit einer geschätzten Gesamtlänge von 3 m ist belegt durch eine weitere Zahnkrone, die durch eine bifurkierende mesiale Carina (genetisch bedingte Abnormität?) bemerkenswert ist. Eine Identifikation ist nicht möglich, aber es zeichnen sich Ähnlichkeiten zu den Coelophysoidea ab. Das Distalende einer Fibula könnte zum gleichen Taxon gehören. Die Dinosaurierfauna der Toutunhe-Formation, bislang auf einen Nodosauriden und nicht näher bestimmtes („megalosaurides“) Theropodenmaterial beschränkt, wird damit um zwei wichtige Komponenten bereichert.


Dinosauria Theropoda Middle Jurassic Junggar Basin Toutnhe Formation 


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  1. Ashraf, R.;Wang, X.-F.;Sun, G.;Li, J. &Mosbrugger, V. 2001. Palynostratigraphic analysis of the Huangshanjie, Hao-jiagou and Badaowan formations in the Junggar Basin (NW China). — In:Sun, G.;Mosbrugger, V.;Ashraf, A.R. &Wang, Y.D., eds., The advanced study of prehistory life and geology of Junggar Basin, Xinjiang, China: 40–64, Urumqi.Google Scholar
  2. Bohlin, B. 1953. Reports from the scientific expedition to the North-Western Provinces of China under leadership of Dr. Sven Hedin. The Sino-Swedish Expedition, Publication 37. VI. Vertebrate Palaeontology. 6. Fossil reptiles of Mongolia and Kansu. — 113 p., Stockholm (Statens Etnografiska Museum).Google Scholar
  3. Charig, A.R. &Milner, A.C. 1990. The systematic position ofBaryonyx walkeri in the light of Gauthier’s reclassification of the Theropoda. — In:Carpenter, K. &Currie, P.J., eds., Dinosaur systematics — approaches and perspectives: 126–140, Cambridge, New York (Cambridge University Press).Google Scholar
  4. Charig, A.R. &Milner, A.C. 1997.Baryonyx walkeri, a fish-eating dinosaur from the Wealden of Surrey. — Bulletin of the Natural History Museum London (Geology)53: 11–70.Google Scholar
  5. Colbert, E.H. 1989. The Triassic dinosaurCoelophysis. — Museum of Northern Arizona Bulletin57: i-xv, 1–160.Google Scholar
  6. Currie, P.j.;Rigby, J.K. &Sloan, R.E. 1990. Theropod teeth from the Judith River Formation of southern Alberta, Canada. — In:Carpenter, K. &Currie, P.J., eds., Dinosaur systematics — approaches and perspectives: 107–125, Cambridge, New York (Cambridge University Press).Google Scholar
  7. Currie, P.J. &Zhao, X.-J. 1993. A new carnosaur (Dinosauria, Theropoda) from the Jurassic of Xinjiang, People’s Republic of China. — Canadian Journal of Earth Sciences30: 2037–2081.Google Scholar
  8. Dong, Z.-M. 1990. On remains of the sauropods from Kelamaili region, Junggar Basin, Xinjiang, China. — Vertebrata PalAsiatica28: 43–58.Google Scholar
  9. Dong, Z.-M. 1993. An ankylosaur (ornithischian dinosaur) from the Middle Jurassic of the Junggar Basin, China. — Vertebrata PalAsiatica31: 257–266.Google Scholar
  10. Dong, Z.-M. 2001. Mesozoic fossil vertebrates from the Junggar Basin and Turpan Basin, Xinjiang, China. — In:Sun, G.;Mosbrugger, V.;Ashraf, A.R. &Wang, Y.D., eds., The advanced study of prehistory life and geology of Junggar Basin, Xinjiang, China: 95–103, Urumqi.Google Scholar
  11. Erickson, G.M. 1995. Split carinae on tyrannosaurid teeth and implications of their development. — Journal of Vertebrate Pale-ontology15: 268–274.Google Scholar
  12. Farlow, J.O.;Brinkman, D.L.;Abler, W.L. &Currie, P.J. 1991. Size, shape and serration density of theropod dinosaur lateral teeth. — Modem Geology16: 161–198.Google Scholar
  13. Gao, Y.-H. 1993. A new species ofSzechuanosaurus from the Middle Jurassic of Dashanpu, Zigong, Sichuan. — Vertebrata PalAsiatica31: 308–314.Google Scholar
  14. Gauthier, J. 1986. Saurischian Monophyly and the Origin of Birds. — In:Padian, K., ed., The origin of birds and the evolution of flight. — Memoirs of the California Academy of Sciences8: 1–55.Google Scholar
  15. Gilmore, C.W. 1920. Osteology of the carnivorous Dinosauria in the United States National Museum, with special reference to the generaAntrodemus (Allosaurus) andCeratosaurus. — Bulletin of the U. S. National Museum110: 1–154.Google Scholar
  16. Hendrix, M.S.;Brassell, S.C.;Carroll, A.R. &Graham, S.A. 1995. Sedimentology, organic geochemistry and petroleum potential of Jurassic Coal Measures: Tarim, Junggar and Turpan basins, Northwest China. — American Association of Petroleum Geologists Bulletin79: 929–959.Google Scholar
  17. Hungerbühler, A. 2000. Heterodonty in the European phytosaurNicrosaurus kapffi and its implications for the taxonomic Utility and functional morphology of phytosaur dentitions. — Journal of Vertebrate Paleontology20: 31–48.CrossRefGoogle Scholar
  18. Hu, S.-J. 1993. A new Theropoda (Dilophosaurus sinensis sp. nov.) from Yunnan, China. — Vertebrata PalAsiatica31: 65–69.Google Scholar
  19. Huene, F. von 1920. Bemerkungen zur Systematik und Stammesgeschichte einiger Reptilien. — Zeitschrift für Induktive Abstammungs- und Vererbungslehre24: 162–166.Google Scholar
  20. Huene, F.von 1932. Die fossile Reptil-Ordnung Saurischia, ihre Entwicklung und Geschichte. — Monographien zur Geologie und Palaeontologie 4 (1–2): viii + 361 p., Berlin, Leipzig.Google Scholar
  21. Huene, F. von 1934. Ein neuer Coelurosaurier in der thüringischen Trias. —Paläontologische Zeitschrift16: 145–170.Google Scholar
  22. Janensch, W. 1925. Die Coelurosaurier und Theropoden der Tendaguru-Schichten Deutsch-Ostafrikas. — Palaeontographica, Supplement-Band7 (1): 1–99.Google Scholar
  23. Madsen, J.H. jr. 1976.Allosaurus fragilis: a revised osteology. — Utah Geological Museum and Mineral Survey Bulletin109: 1–163.Google Scholar
  24. Maisch, M.W.;Matzke, A.T.;Pfretzschner, H.-U.;Ye, J. &Sun, G. 2001. The fossil vertebrate faunas of the Toutunhe and Qigu Formations of the Southern Junggar Basin and their biostratiugraphical and palecological implications. — In:Sun, G.;Mosbrugger, V.;Ashraf, A.R. &Wang, Y.D., eds., The advanced study of prehistory life and geology of Junggar Basin, Xinjiang, China: 83–94, Urumqi.Google Scholar
  25. Maisch, M.W.;Matzke, A.T.;Pfretzschner, H.-U.;Sun, G.;Stöhr, H. &Grossmann, F. in press. Fossil vertebrates from the Middle and Upper Jurassic of the Southern Junggar Basin (NW China) — results of the Sino-German Expeditions 1999–2000. — Neues Jahrbuch für Geologie und Paläontologie Abhandlungen.Google Scholar
  26. Marsh, O.C. 1881. Principal characters of American dinosaurs. Part 5. — American Journal of Science (3)23: 81–86.Google Scholar
  27. Marsh, O.C. 1884. Principal characters of American Jurassic dinosaurs. Part 8. The Order Theropoda. — American Journal of Science (3)27: 329–340.Google Scholar
  28. McKnight, C.L.;Graham, S.A.;Carroll, A.R.;Gan, Q.;Dilcher, D.L.;Zhao, M. &Liang, Y.H. 1990. Fluvial sedimentology of an Upper Jurassic petrified forest assemblage, Shishu Formation, Junggar Basin, Xinjiang, China. — Paleogeography, Paleoecology, Paleoclimatology79: 1–9.CrossRefGoogle Scholar
  29. Meyer, H. von 1861. Reptilien aus dem Stubensandstein des oberen Keupers. — Palaeontographica10: 227–246.Google Scholar
  30. Owen, R. 1842. Report on British fossil reptiles. Part 2. — Report of the British Association for the Advancement of Science11 [1841]: 60–294.Google Scholar
  31. Pfretzschner, H.U.;Ashraf, A.R.;Maisch, M.W.;Sun, G.;Wang, Y.D. &Mosbrugger, V. 2001. Cyclic growth in dinosaur bones from the Upper Jurassic of NW China and its paleoclimatic implications. — In:Sun, G.;Mosbrugger, V.;Ashraf, A.R. &Wang, Y.D., eds., The advanced study of prehistory life and geology of Junggar Basin, Xinjiang, China: 21–39, Urumqi.Google Scholar
  32. Raath, M. 1990. Morphological variation in small theropods and its meaning in systematics: Evidence fromSyntarsus rhodesiensis. — In:Carpenter, K. &Currie, P.J., eds., Dinosaur systematics — approaches and perspectives: 91–105, Cambridge, New York (Cambridge University Press).Google Scholar
  33. Rauhut, O.W.M. &Kriwet, J. 1994. Teeth of a big theropod dinosaur from Porto das Barcas, Portugal. — Berliner Geowissenschaftliche Abhandlungen (E)13: 179–185.Google Scholar
  34. Rowe, T. 1989. A new species of the theropod dinosaurSyntarsus from the Early Jurassic Kayenta Formation of Arizona. — Journal of Vertebrate Paleontology9: 125–136.Google Scholar
  35. Russell, D.A. &Zheng, Z. 1993. A new large mamenchisaurid from the Junggar Basin, Xinjiang, People’s Republic of China. — Canadian Journal of Earth Sciences30: 2082–2095.Google Scholar
  36. Seeley, H.G. 1887. The classification of the Dinosauria. — Report of the British Association for the Advancement of Science57 [1887]: 698–699.Google Scholar
  37. Wang. Y.-D.;Zhang, W. &Saki, K. 2000. Fossil woods from the Upper Jurassic of Qitai, Junggar Basin, Xinjiang, China. — Acta Palaeontologica Sinica39: 176–185.Google Scholar
  38. Welles, S.P. 1984.Dilophosaurus wetherilli (Dinosauria, Theropoda) osteology and comparisons. — Palaeontographica (A)185: 85–180.Google Scholar
  39. Zhao, X.-J. 1993. A new Mid-Jurassic sauropod(Klamelisaurus gobiensis gen. et sp. nov.) from Xinjiang, China. — Vertebrata PalAsiatica31: 132–138.Google Scholar
  40. Zhao, X.-J. &Currie, P.J. 1993. A large crested theropod from the Jurassic of Xinjiang, People’s Republic of China. — Canadian Journal of Earth Sciences30 (10/11): 2027–2036.Google Scholar

Copyright information

© E. Schweizerbart’sche Verlagsbuchhandlung 2003

Authors and Affiliations

  1. 1.Institut und Museum für Geologie und Paläontologie der Eberhard-Karls-Universität TübingenTübingenGermany

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