Anisodon (Perissodactyla, Chalicotheriinae) from the middle Miocene locality Gračanica (Bugojno Basin, Gornji Vakuf, Bosnia and Herzegovina)

  • Margery C. CoombsEmail author
  • Ursula B. Göhlich
Original Paper


The fossil mammal fauna of the Gračanica coal mine in central Bosnia and Herzegovina includes dental remains of Anisodon (Perissodactyla, Chalicotheriidae, Chalicotheriinae). These teeth most closely resemble those of Anisodon grande from Sansan (France, MN6) and Neudorf an der March (Slovakia, MN6), and are tentatively referred to that species. Anisodon is first known in Europe from the Loire Basin (France), which has been dated as MN5. Given that the Gračanica fauna is considered to be MN5 or earliest MN6 (Langhian), the Gračanica chalicothere represents one of the earlier known finds of both the Chalicotheriinae and Anisodon in Europe. The first known members of the Chalicotheriinae are from the earliest Miocene of Asia. By the middle Miocene, Anisodon had appeared and is known from Asia and Europe, to which it had recently immigrated.


Anisodon grande Chalicotheriinae Gračanica Miocene Early Langhian 



Lilith-Valerie Svaricek (University Vienna) assisted in picture editing. We are grateful for helpful review comments by Denis Geraads (Muséum National d’Histoire Naturelle, Paris) and Kurt Heissig (Bayerische Staatssammlung für Paläontologie und Geologie, Munich).

Compliance with ethical standards

Conflict of interest

The authors declare that they have no conflict of interest.


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Authors and Affiliations

  1. 1.Department of BiologyUniversity of MassachusettsAmherstUSA
  2. 2.Department of Geology and PaleontologyNatural History Museum ViennaViennaAustria

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