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New trepostome Bryozoa from the Early Triassic (Smithian/Spathian) of Nevada

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Zwei neue Bryozoen-Arten der Familie Dyscritellidae (Trepostomata),Dyscritellopsis thaynesianus n. sp. undDyscritellopsis montelloensis n. sp., werden aus den Thaynes Kalken der unteren Trias (Smithian/Spathian) von Nevada (USA) beschrieben. Die Bryozoen-Fauna liefert weiteren Beleg für das Überleben paläozoischer Stammlinien in frühest triadischer Zeit auf den offenen Schelfen der Nordhemisphere außerhalb der Tropen. Sie zeigt paläobiogeo-graphische Beziehungen zu Faunen der Unteren Trias von Spitsbergen.


Two new bryozoan species of the trepostome family Dyscritellidae,Dyscritellopsis thaynesianus n. sp. andDyscritellopsis montelloensis n. sp., are described from the Early Triassic (Smithian/Spathian) Thaynes Limestone, Nevada (USA). The bryozoan fauna documents the survival of Paleozoic lineages into the earliest Triassic on northern open shelves outside the tropics. The fauna holds paleobiogeo-graphic connections to the Early Triassic bryozoan faunas of Spitsbergen.

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  1. Astrova, G.G. 1965. The morphology, evolution, and systematics of the Ordovician and Silurian Bryozoa. — Trudy Paleontological Institute, Akademia Nauk SSR106: 1–431.

  2. Astrova, G.G. 1978. The history of development, systematics, and phylogeny of Bryozoa: Order Trepostomata — Trudy Paleontological Institute, Akademia Nauk SSR169: 1–240.

  3. Bissell, H.J. 1973. Permian-Triassic boundary in the Eastern Great Basin area. — In:Logan, A. &Hills, L.V., eds., The Permian and Triassic systems and their mutual boundary: 700–726, Calgary (Canadian Society of Petroleum Geologists).

  4. Bolton, T.E. 1961. AdditionalArcticopora from the Triassic of northeastern Ellesmere Island. — Proceedings of the Geological Association of Canada13: 55–56.

  5. Borg, F. 1926. Studies on Recent cyclostomatous Bryozoa. — Zoologiska Bidrag frå Uppsala10: 181–507.

  6. Carr, T.R. &Paull, R.K. 1983. Early Triassic stratigraphy and paleogeography of the Cordilleran Miosyncline. — In:Reynolds, M.W. &Dolly, E.D., eds., Mesozoic paleogeography of the west-central Unites States: 339–355, Denver (Rocky Mountains Section S.E.P.M.).

  7. Clark, D.L. 1957. Marine Triassic stratigraphy in eastern Great Basin. — American Association of Petroleum Geologists, Bulletin41 (10): 2192–2222.

  8. Collinson, J.W.;Kendall, C.G.S.C. &Marcantel, J.B. 1976. Permian-Triassic boundary in eastern Nevada and west-central Utah. — Geological Society of America, Bulletin87: 821–824.

  9. Dunaeva, N.N. &Morozova, LP. 1967: Evolutionary features and the systematic position of some late Paleozoic trepostomes. — Paleontological Zhurnal1967 (4): 86–94.

  10. Duncan, H. 1939. Trepostomatous Bryozoa from the Traverse Group of Michigan. — Contributions of the Museum of Paleontology of the University of Michigan5 (10): 171–270.

  11. Ehrenberg, C.G. 1831. Symbolae physicae, seu icones et descriptiones corporum naturalium novorum aut minus cognitorum, quae ex itineribus per Libyam, Aegyptum, Nubium, Dongalam, Syriam, Arabiam et Habessiniam… studio annis 1820–25 redierunt… Pars Zoologica, 4. Animalia Evertebrata exclusis Insectis. — 10 pis. (Berolini).

  12. Engeser, F.S. &Taylor, P.D. 1991. Supposed Triassic bryozoans in the Klippstein collection from the Italian Dolomites rede-scribed as calcified demosponges. — Bulletin of the British Museum of Natural History45 (1): 39–55.

  13. Ernst, A. 2001, Bryozoa from the Upper Permian Zechstein Formation of Germany. — Senckenbergiana lethaea81 (1): 135- 181.

  14. Ernst, A. 2002. Systematics and biogeography of the Permian bryozoans in Europe. — In:Wyse Jackson. P.N.;Buttler, C. &Spencer Jones, M.E., eds., Bryozoan Studies 2001: 109–112, Lisse (Balkema Publishers).

  15. Erwin, D.H. 1994. The Permo-Triassic extinction. — Nature367: 231–236.

  16. Flügel, E. 1994. Pangean shelf carbonates: Controls and paleoclimatic significance of Permian and Triassic reefs. — Geological Society of America, Special Paper288: 247–266.

  17. Flügel, E. &Reinhardt, J.w. 1989. Uppermost Permian reefs in Skyros (Greece) and Sichuan (China): Implications for the Late Permian extinction event. — Palaios4 (6): 502–518.

  18. Flügel, E. &Senowbari-Daryan, B. 1996. Evolution of Triassic reef biota: State of the art. — Göttinger Arbeiten zur Geologie und PaläontologieSB2: 285–294.

  19. Flügel, E. &Senowbari-Daryan, B. 2001: Triassic reefs of the Tethys. — In:Stanley, G.D., jr., ed., The history and sedimentology of ancient reef systems: 217–249, New York (Kluwer Academic/Plenum Publishers).

  20. Fois, E. &Gaetani, M. 1984. The recovery of reef-building communities and the role of cnidarians in carbonate sequences of the Middle Triassic (Anisian) in the Italian Dolomites. — Palaeontographica Americana54: 191–200.

  21. Fritz, M. 1961. A new bryozoan genus from Lake Hazen, Northeastern Ellesmere Island. — Proceedings of the Geological Association of Canada13: 53–55.

  22. Gilmour, E.H. &Morozova, LP. 1999. Biogeography of the Late Permian bryozoans. — Paleontological Zhurnal33 (1): 36–51 (English translation).

  23. Girty, G.H. 1911. New genera and species of Carboniferous fossils from the Fayetteville shale of Arkansas. — Annals of the New York Academy of Science20 (3): 189–238.

  24. Girty, G.H. 1920. Carboniferous and Triassic faunas. — United States Geological Survey, Professional Paper111: 641–657.

  25. Henrich, R.;Freiwald, A.;Schäfer, P.;Wehrmann, A.;Kühlmann, D.;Brachert, T.;Betzler, C;Samtleben, C;Bader, B. &Zankl, H. 1995. Controls of reef formation and biogenic carbonate sedimentation in non-tropical environmental settings: modern case studies and fossil counterparts. — Facies32: 71–108.

  26. James, N.P. 1997. The cool-water carbonate depositional realm. — Society of Economic Paleontologists and Mineralogists (S.E.P.M.), Special Publication56: 1–20.

  27. Lazutktna, O.F. 1963. Discoveries of bryozoans of the Paleozoic genusBatostomella in the Triassic system. — Paleontological Zhurnal1963 (4): 126–128 (in Russian).

  28. Manfield, G.R. 1927. Geography, geology, and mineral resources of part of southeastern Idaho. — United States Geological Survey, Professional Paper152: 1–453.

  29. Morozova, L.P. 1960. The genusPseudobatostomella gen. nov. — In: Fundamentals of Paleontology. Bryozoa and Brachiopoda: 1–71, Moscow (in Russian).

  30. Morozova, L.P. 1965. Bryozoa. — In:Ruzhentev, V.E., ed., The Evolution and Changes in the Marine Organisms at the Paleozoic-Mesozoic Boundary: 57–62, 183–197, Moscow (in Russian).

  31. Morozova, L.P. 1969. Systematic composition and range of bryozoans in the Triassic. — Paleontological Zhurnal1969 (2): 49–57 (in Russian).

  32. Morozova, LP. 1970. Late Permian Bryozoa. — Trudy Paleontological Institute122: 1–347 (in Russian).

  33. Morozova, L.P. &Kruchinina, O.N. 1986. The Permian bryozoans of the Arctic Region (western Sector). — Akademia Nauk, Moscow (English translation).

  34. Nakrem, H.A. 1994. Middle Carboniferous to Early Permian bryozoans from Spitsbergen. — Acta Palaeontologica Polonica39 (1): 45–116.

  35. Nakrem, H.A. &Mørk, A. 1991. New Early Triassic Bryozoa (Trepostomata) from Spitsbergen, with some remarks on the stratigraphy of the investigated horizons. — Geological Magazine128 (2): 129–140.

  36. Nelson, CS. 1988. An introductory perspective on non-tropical shelf carbonates. — Sedimentary Geology60: 3–12.

  37. Nikiforova, A.I. 1933. Upper Paleozoic bryozoans of Julfinskiy region. — Trudy Vsesojuznogo geologo-razvedochnogo objedinenija364: 1–44.

  38. Paull, R.K. &Paull, R.A. 1986. Epilogue for the Permian in the western Cordillera — a retrospective view from the Triassic. — Contributions of Geology, University of Wyoming24: 243–252.

  39. Rao, C.P. 1981. Criteria for the recognition of cold-water sedimentation: Berridale Limestone (Lower Permian), Tasmania, Australia. — Journal of Sedimentary Petrology51 (2): 491–506.

  40. Redfern, R. 1986. The making of a continent. — 241 p., New York (Times Books).

  41. Ross, J. 1978. Biogeography of Permian ectoproct Bryozoa. — Palaeontology21 (2): 341–356.

  42. Sakagami, S. 1975. Permian Bryozoa from Khao Hin Kling, near Phetchabun, North-Central Thailand. — Geology and Palaeontology of Southeast Asia16: 33–43.

  43. Sakagami, S. 1998. Permian Bryozoans from North Ellesmere Island, Canadian Arctic Archipelago. — Bulletin of the Natural Science Museum, Tokyo (C)24 (2): 67–91.

  44. Schäfer, P. 1994. Bryozoen der Trias — eine Übersicht. — Abhandlungen der Geologischen Bundes-Anstalt Wien50: 387–397.

  45. Schäfer, P. &Fois, E. 1987. Systematics and evolution of Triassic Bryozoa. — Geologica et Palaeontologica21: 173–225.

  46. Schäfer, P. &Grant-Mackie, J.-A. 1994. Triassic Bryozoa from the Murihiku and Toriesse Supergroups, New Zealand. — Memoirs of the Association of Australasian Palaeontologists16: 1–52.

  47. Schäfer, P. &Grant-Mackie, J.-A. 1998. Revised systematics and palaeobiogeography of some Late Triassic colonial invertebrates from the Pacific region. — Alcheringa22: 87–122.

  48. Schäfer, P.;Senowbari-Daryan, B. &Hamedani, A. 2003. Stenolaemate bryozoans from the Upper Triassic (Norian-Rhaetian) Nayband Formation near Wali Abad, SE Abadeh (Central Iran). — Facies48: 135–150.

  49. Schubert, J.K. &Bottjer D.J. 1995. Aftermath of the Permian-Triassic mass extinction event: Paleoecology of Lower Triassic carbonates in the western USA. — Palaeogeography, Palaeoclimatology, Palaeoecology116 (1995): 1–39.

  50. Snyder, E.M. 1976. Taxonomy and biostratigraphy of the Bryozoa of the Gerster Formation (Permian), northeastern Nevada. — Unpublished Ph. D. thesis, Eastern Washington University. -171 p., Cheney, Wash.

  51. Taylor, P.D. &Michalik, J. 1991. Cyclostome bryozoans from the Late Triassic (Rhaetian) of the West Carpathians, Czechoslovakia. — Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen182 (3): 285–302.

  52. Teichert, C. 1990. The Permian-Triassic boundary revisited. — In:Kauffman, E.G. &Walliser, O.H., eds., Extinction events in earth history: 199–238, Heidelberg-Berlin (Spinger).

  53. Teichert, C. &Kümmel, B. 1976. Permian-Triassic boundary in the Kap Stosch area, East Greenland. — Meddelser om Grønland197 (5): 1–49.

  54. Troitskaya, T.D. 1970. Some Famennian bryozoans from Central Kazakhstan (Sarysu-Teniz watershed) — In:Astrova, G.G. &Chudinova, LI., eds., New species of Paleozoic Bryozoa and corals: 40–46, Moscow (Nauka Press).

  55. Trümpy, R. 1960. Über die Perm-Trias-Grenze in Ostgrönland und die Problematik stratigraphischer Grenzen. — Geologische Rundschau49: 97–111.

  56. Ulrich, E.O. 1882. American Paleozoic Bryozoa. — Journal of the Cincinnati Society of Natural History5: 121–175.

  57. Valentine, J.W. &Moore, E.M. 1973. Provincialism and diversity across the Permo-Triassic boundary. — In:Logan, A. &Hills, L.V., eds., The Permian and Triassic systems and their mutual boundary: 759–766, Calgary (Canadian Society of Petroleum Geologists).

  58. Waagen, W. &Wentzel, J. 1886. Coelenterata. Salt Lake fossils. — Palaeontographica Indica13 (1): 854–892, 904–924, 963–966.

  59. Wass, R.E. 1968. Permian Polyzoa from the Bowen Basin, Queensland. — Bulletin of the Bureau of Mineral Resources, Geology and Geophysics of Australia90: 1–134.

  60. WWW.SCOTESE.COM/climate.htm

  61. Yang, K. &Hsia, F.S. 1975. Palaeontology. — Reports of scientific expedition in the Mount Jolmo Lungma region (1966–1968)1: 39–57.

  62. Yang, F.-Z. &Lu, L.-H. 1981. Geographical and stratigraphical distributions of Permian Bryozoa in China. — In:Larwood, G.P. &Nielsen, C, eds., Recent and fossil Bryozoa: 305–307, Fredensborg (Olsen & Olsen).

  63. Young, A.R. 1998. Geology of the Goose Creek Mountains, northwestern Box Elder County, Utah, and northeastern Elko County, Nevada. — Unpublished Ph. D. thesis, University of Utah, Salt Lake City, Utah.

  64. Young, J. 1883. On Ure’s“Millepora”, Tabulipora (Cellepord) urei Fleming. — Annals and Magazine of Natural History5 (12): 1–158.

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Correspondence to Priska Schäfer.

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Schäfer, P., Cuffey, R.J. & Albert, R.Y. New trepostome Bryozoa from the Early Triassic (Smithian/Spathian) of Nevada. Paläontol Z 77, 323–340 (2003). https://doi.org/10.1007/BF03006945

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  • Trepostomata
  • Smithium/Spathium
  • Nordost Nevada
  • Thaynes Kalke
  • Systematik
  • Paläobiogeographie
  • Überlebende


  • Trepostomata
  • Smithian/Spathian
  • Northeast Nevada
  • Thaynes Limestone
  • systematics
  • paleobiogeography
  • mass extinction survivors.